rod << bk1 << Med << bk2
Dominantly enhances position effect variegation at the w locus caused by In(1)wm4, C(1;YL)wmMc or T(1;4)wm5 and position effect variegation at the y locus caused by In(1)y3P. Position effect variegation at the w locus is not seen when Df(3R)E40 is combined with In(1)wm4rv5, In(1)wm4rv6 or In(1)wm4rv26.
Deficient embryos show a variably penetrant mutant midgut phenotype: central constriction absent.
Homozygous embryos do not complete head involution or dorsal closure, and tracheae are not connected. Midgut constrictions are incomplete, Malpighian tubules have a variable diameter along their length and the hindgut is sometimes shorter than normal.
Strictly maternal enhancer of dpphr4 lethality. The lethal interaction is due to a loss of dorsal-most fates in the embryos, demonstrated by loss of amnioserosa cells.