About 3% of Dllmd23/Dll9 animals emerge as adults, with all escapers displaying partial transformation of the antenna to leg structures and deletion of distal leg structures.
Homozygote has a very mild leg phenotype: males have a decrease in number of sex comb teeth. Entire arista and distal most portion of AIII are transformed into a distally complete tarsus.
Minor reductions in the tarsal region.
Heterozygotes are wild-type. Homozygotes show a transformation of antenna to leg. Hemizygous embryos but not homozygous embryos lack the Keilin's organs, and sometimes lack the labial sense organ, antennal sense organ and maxillary sense organ.
Homozygotes show partial transformation of antennae to legs as well as deletions of some leg structures. Tarsal tissue, sometimes including claws, develops in place of arista and part of third antennal segment. Third segment usually contains patchwork of incompletely differentiated leg tissue; first and second segments normal. Leg effects distal to mid-tibia; legs proximal to mid-tibia normal. In extreme cases number of tarsal segments reduced to two or three; basitarsus and distal tibia may be missing; in most extreme cases claws may also be missing. In the case of mild expression abnormal bristle patterns, including polarity reversal, occur around basitarsus-tibia joint, where there may also develop cuticular hyperplasia. Penetrance greater in females than males, in antennae than legs and in metathoracic than other legs.