Allele Dmel\fz²¹

General Information
Symbol	Dmel\fz²¹	Species	D. melanogaster
Name		FlyBase ID	FBal0004937
Feature type	allele	Associated gene	Dmel\fz
Also Known As	fz^P21

Allele class
Mutagen	ethyl methanesulfonate
Recent Updates
Description	What does this section display? This section contains items that were added to this record for each release. It currently only tracks new links between this FlyBase report and other FlyBase data classes (e.g. genes, references, stocks) or controlled vocabulary terms (e.g. GO, anatomy terms). What does this section not display? This section does not currently display links that were removed or gene model changes.
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FB2013_03
FB2013_02
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Nature of the Allele
Allele class
Mutagen	ethyl methanesulfonate
Mutations Mapped to the Genome

Type Location Additional Notes References
Associated Sequence Data
DDBJ / EMBL / GenBank	DNA sequence Protein sequence Name

UniProtKB/Swiss-Prot
UniProtKB/TrEMBL

Progenitor genotype
Nature of the lesion	Statement Reference Insertion in the cysteine-rich domain (CRD) that results in a truncated protein being produced. (Povelones et al., 2005) Insertion into the first exon that shifts the reading frame. (Jones et al., 1996)
Caused by insertion	?{}fz²¹ (Povelones et al., 2005, Weber et al., 2012)
Cytology
Phenotypic Data
Phenotypic Class
	planar polarity defective (Olguín et al., 2011) planar polarity defective, with Scer\GAL4^hh-Gal4, fz^Scer\UAS.cSa (Lawrence et al., 2004) planar polarity defective (with fz¹⁹) (Weber et al., 2008) planar polarity defective (with fz²³) (Wu et al., 2004) planar polarity defective \| cell non-autonomous (Adler et al., 2000) planar polarity defective \| cell non-autonomous \| somatic clone (Lee and Adler, 2002, Olguín et al., 2011) planar polarity defective \| recessive (Pataki et al., 2010) planar polarity defective \| somatic clone (Collier et al., 2005) visible visible \| recessive (Pataki et al., 2010) visible \| somatic clone (Collier et al., 2005)
Phenotype Manifest In
	dorsal wing blade (with fz²³) (Doyle et al., 2008) embryonic/larval dorsal trunk (with fz³⁰) (Chung et al., 2009) eye (with fz²³) (Wu et al., 2004) macrochaeta & adult thorax ommatidium (with fz¹⁹) (Weber et al., 2008) trichome & pleural membrane, with Scer\GAL4^hh-Gal4, fz^Scer\UAS.cSa (Lawrence et al., 2004) wing (Jones et al., 1996) wing \| cell non-autonomous \| somatic clone (Adler et al., 2000) wing cell \| P-stage (Classen et al., 2005) wing hair (Pataki et al., 2010) wing hair (with fz²³) (Wu et al., 2004, Wu et al., 2008) wing hair \| cell non-autonomous \| somatic clone (Lee and Adler, 2002) wing hair \| somatic clone (Collier et al., 2005)
Detailed Description
	Statement Reference The orientation of trichomes is uniformly randomised only in the most central region of the notum in mutant animals. (Olguín et al., 2011) Homozygous clones in the notum nonautonomously influence the polarity of neighbouring bristles and trichomes. (Olguín et al., 2011) fz[21] mutant wings display strong hair orientation defects, but very few if any multiple hairs. (Pataki et al., 2010) fz[21]/fz[30] stage 16 embryos have longer dorsal trunks than normal. (Chung et al., 2009) In contrast to wild-types, no clear wing membrane ridge orientation is apparent in either the anterior or posterior region of the wing of fz[23]/fz[21] mutants. (Doyle et al., 2008) Prehair initiation is delayed in mutant cells away from the clone edge in homozygous clones in the wing. (Strutt and Warrington, 2008) Approximately 16% of ommatidia are symmetrical in fz[19]/fz[21] flies. (Weber et al., 2008) The wing hairs in fz[21]/fz[23] mutants no longer point uniformly from proximal to distal as is seen in wild type. (Wu et al., 2008) Most larvae from fz¹⁵/fz²¹ mothers crossed with fz¹⁵/+ fathers have cuticles with normal morphology; only 2 of 30 exhibit fusions between segments. Another 2 mutants exhibit misplaced actin-based protrusions in denticle row 1. (Walters et al., 2006) The hexagonal packing of intervein cells, which usually occurs between wing development stage P2B (when the first morphological signs of veins appear (FBrf0005070), and the middle of P2C (before hair formation (FBrf0005070)) is partially disrupted in fz²¹/fz²¹ flies. (Classen et al., 2005) Mutant clones in the wing result in domineering nonautonomy wing hair polarity phenotype. (Collier et al., 2005) Somatic clones in the wing disc do not exhibit significant delay in hair development. (Lee and Adler, 2004) fz¹⁵/fz²¹ ovaries show only very minor abnormalities. (Cohen et al., 2002) Mutant clones in the wing show domineering non-autonomy: cells distal to the clone, as well as within the clone, show alteration of polarity of wing hairs. (Lee and Adler, 2002) Homozygous somatic clones in the wing cause domineering non-autonomy. (Adler et al., 2000) Dendritic development in fz²¹/Df(3L)fz-D21 mutants is indistinguishable from wild-type. (Gao et al., 2000) Non-autonomous phenotype in wing clones. (Jones et al., 1996) Moderate thoracic bristle phenotype; moderate wing-hair disorientation.
External Data
Linkouts
Interactions
	Show the genetic interaction network for Enhancers & Suppressors
Phenotypic Class
Enhanced by
Statement Reference fz²¹ has visible \| recessive phenotype, enhanceable by Rab23[+]/Rab23⁵¹ (Pataki et al., 2010) fz²¹ has visible \| recessive phenotype, enhanceable by Rab23⁵¹ (Pataki et al., 2010)
NOT Enhanced by
Statement Reference fz²¹ has planar polarity defective phenotype, non-enhanceable by pk^pk-sple-13/pk^pk-sple-13 (Lawrence et al., 2004) fz²¹ has planar polarity defective phenotype, non-enhanceable by pk^{sple.Scer\UAS}/Scer\GAL4^hh-Gal4 (Lawrence et al., 2004)
Suppressed by
Statement Reference fz²¹/fz¹⁹ has planar polarity defective phenotype, suppressible by Egfr[+]/Egfr^top-18A (Weber et al., 2008) fz²¹ has planar polarity defective \| somatic clone \| cell non-autonomous phenotype, suppressible by in^IH56/in^unspecified (Lee and Adler, 2002) fz²³/fz²¹ has planar polarity defective phenotype, suppressible by fz::fz2^1-1-2.tub.W (Wu et al., 2004) fz²³/fz²¹ has planar polarity defective phenotype, suppressible by fz::fz2^1-1-2S.tub.W (Wu et al., 2004) fz²³/fz²¹ has planar polarity defective phenotype, suppressible by fz::fz2^1-2-1.tub.W (Wu et al., 2004)
NOT suppressed by
Statement Reference fz²¹ has planar polarity defective \| somatic clone \| cell autonomous phenotype, non-suppressible by in^IH56/in^unspecified (Lee and Adler, 2002) fz²¹ has planar polarity defective \| somatic clone \| cell non-autonomous phenotype, non-suppressible by in^IH56 (Lee and Adler, 2002) fz²¹ has planar polarity defective \| somatic clone phenotype, non-suppressible by Vang^stbm-6 (Wu and Mlodzik, 2008) fz²¹ has planar polarity defective phenotype, non-suppressible by pk^pk-sple-13/pk^pk-sple-13 (Lawrence et al., 2004) fz²¹ has planar polarity defective phenotype, non-suppressible by pk^{sple.Scer\UAS}/Scer\GAL4^hh-Gal4 (Lawrence et al., 2004) fz²³/fz²¹ has planar polarity defective \| somatic clone phenotype, non-suppressible by fz::fz2^{2-1-1.tub.T:Hsap\MYC} (Wu and Mlodzik, 2008) fz²³/fz²¹ has planar polarity defective phenotype, non-suppressible by fz::fz2^1-2-2.tub.W (Wu et al., 2004) fz²³/fz²¹ has planar polarity defective phenotype, non-suppressible by fz::fz2^2-1-1.tub.W (Wu et al., 2004)
Enhancer of
Statement Reference fz²¹ is an enhancer of planar polarity defective \| recessive phenotype of Rab23⁵¹ (Pataki et al., 2010)
NOT Enhancer of
Statement Reference fz[+]/fz²¹ is a non-enhancer of visible phenotype of Scer\GAL4^en-e16E, kermit^GS2053 (Djiane and Mlodzik, 2010)
Suppressor of
Statement Reference fz[+]/fz²¹ is a suppressor \| partially of planar polarity defective \| dominant phenotype of Egfr^f24 (Weber et al., 2008) fz[+]/fz²¹ is a suppressor \| partially of planar polarity defective phenotype of Egfr^t1/Egfr^f11 (Weber et al., 2008) fz²¹, fz³⁷ is a suppressor of planar polarity defective phenotype of Scer\GAL4^ptc-559.1, fz³⁷ (Lawrence et al., 2004) fz²¹/fz²¹ is a suppressor of planar polarity defective \| somatic clone \| cell non-autonomous phenotype of Scer\GAL4^αTub84B.PL, pwn^unspecified, stan^Scer\UAS.cUa (Lawrence et al., 2004, Lawrence et al., 2004) fz²¹/fz²¹ is a suppressor of planar polarity defective \| somatic clone phenotype of Scer\GAL4^αTub84B.PL, pwn^unspecified, stan^Scer\UAS.cUa (Lawrence et al., 2004, Lawrence et al., 2004) fz²¹ is a suppressor of planar polarity defective \| somatic clone phenotype of Vang^stbm-6 (Wu and Mlodzik, 2008) fz³⁷/fz²¹ is a suppressor of planar polarity defective phenotype of Scer\GAL4^ptc-559.1, stan^Scer\UAS.cUa (Lawrence et al., 2004)
NOT Suppressor of
Statement Reference fz[+]/fz²¹ is a non-suppressor of visible phenotype of Scer\GAL4^en-e16E, kermit^GS2053 (Djiane and Mlodzik, 2010)
Other
Statement Reference chas¹, fz²¹ has planar polarity defective phenotype (Olguín et al., 2011)
Phenotype Manifest In
Enhanced by
Statement Reference fz²¹ has wing hair phenotype, enhanceable by Rab23[+]/Rab23⁵¹ (Pataki et al., 2010) fz²¹ has wing hair phenotype, enhanceable by Rab23⁵¹ (Pataki et al., 2010)
NOT Enhanced by
Statement Reference fz²¹ has trichome & adult abdomen phenotype, non-enhanceable by pk^pk-sple-13/pk^pk-sple-13 (Lawrence et al., 2004) fz²¹ has trichome & adult abdomen phenotype, non-enhanceable by pk^{sple.Scer\UAS}/Scer\GAL4^hh-Gal4 (Lawrence et al., 2004)
Suppressed by
Statement Reference fz²¹/fz¹⁹ has ommatidium phenotype, suppressible by Egfr[+]/Egfr^top-18A (Weber et al., 2008) fz²¹ has wing hair \| somatic clone \| cell non-autonomous phenotype, suppressible by in^IH56/in^unspecified (Lee and Adler, 2002) fz²³/fz²¹ has eye phenotype, suppressible by fz::fz2^1-1-2.tub.W (Wu et al., 2004) fz²³/fz²¹ has eye phenotype, suppressible by fz::fz2^1-1-2S.tub.W (Wu et al., 2004) fz²³/fz²¹ has eye phenotype, suppressible by fz::fz2^1-2-1.tub.W (Wu et al., 2004) fz²³/fz²¹ has wing hair phenotype, suppressible by fz::fz2^1-1-2.tub.W (Wu et al., 2004) fz²³/fz²¹ has wing hair phenotype, suppressible by fz::fz2^1-1-2S.tub.W (Wu et al., 2004) fz²³/fz²¹ has wing hair phenotype, suppressible by fz::fz2^1-2-1.tub.W (Wu et al., 2004)
NOT suppressed by
Statement Reference fz²¹ has trichome & adult abdomen phenotype, non-suppressible by pk^pk-sple-13/pk^pk-sple-13 (Lawrence et al., 2004) fz²¹ has trichome & adult abdomen phenotype, non-suppressible by pk^{sple.Scer\UAS}/Scer\GAL4^hh-Gal4 (Lawrence et al., 2004) fz²¹ has wing hair \| somatic clone \| cell autonomous phenotype, non-suppressible by in^IH56/in^unspecified (Lee and Adler, 2002) fz²¹ has wing hair \| somatic clone \| cell non-autonomous phenotype, non-suppressible by in^IH56 (Lee and Adler, 2002) fz²¹ has wing hair \| somatic clone phenotype, non-suppressible by Vang^stbm-6 (Wu and Mlodzik, 2008) fz²³/fz²¹ has eye phenotype, non-suppressible by fz::fz2^1-2-2.tub.W (Wu et al., 2004) fz²³/fz²¹ has eye phenotype, non-suppressible by fz::fz2^2-1-1.tub.W (Wu et al., 2004) fz²³/fz²¹ has wing hair \| somatic clone phenotype, non-suppressible by fz::fz2^{2-1-1.tub.T:Hsap\MYC} (Wu and Mlodzik, 2008) fz²³/fz²¹ has wing hair phenotype, non-suppressible by fz::fz2^1-2-2.tub.W (Wu et al., 2004) fz²³/fz²¹ has wing hair phenotype, non-suppressible by fz::fz2^2-1-1.tub.W (Wu et al., 2004)
NOT Enhancer of
Statement Reference fz[+]/fz²¹ is a non-enhancer of wing hair phenotype of Scer\GAL4^en-e16E, kermit^GS2053 (Djiane and Mlodzik, 2010)
Suppressor of
Statement Reference fz[+]/fz²¹ is a suppressor \| partially of ommatidium phenotype of Egfr^f24 (Weber et al., 2008) fz[+]/fz²¹ is a suppressor \| partially of ommatidium phenotype of Egfr^t1/Egfr^f11 (Weber et al., 2008) fz²¹, fz³⁷ is a suppressor of trichome & adult abdomen phenotype of Scer\GAL4^ptc-559.1, fz³⁷ (Lawrence et al., 2004) fz²¹/fz²¹ is a suppressor of trichome & adult abdomen \| somatic clone \| cell non-autonomous phenotype of Vang^stbm-6 (Lawrence et al., 2004, Lawrence et al., 2004) fz²¹ is a suppressor of wing hair \| somatic clone phenotype of Vang^stbm-6 (Wu and Mlodzik, 2008)
NOT Suppressor of
Statement Reference fz[+]/fz²¹ is a non-suppressor of wing hair phenotype of Scer\GAL4^en-e16E, kermit^GS2053 (Djiane and Mlodzik, 2010)
Other
Statement Reference chas¹, fz²¹ has chaeta phenotype (Olguín et al., 2011) Df(3L)fz2, fz²¹ has cardioblast phenotype (Bhanot et al., 1999) Df(3L)fz2, fz²¹ has midgut constriction phenotype (Bhanot et al., 1999) Df(3L)fz2, fz²¹ has RP2 motor neuron phenotype (Bhanot et al., 1999) Df(3L)fz2/+, fz²¹ has embryonic/first instar larval cuticle phenotype (Chen et al., 2004) fz2^C2, fz²¹ has wing margin \| somatic clone phenotype (Chen et al., 2004, Chen et al., 2004) fz2^C2, fz²¹ has wing margin bristle \| somatic clone phenotype (Chen et al., 2004, Chen et al., 2004)
Additional Comments
Genetic Interactions
Statement Reference chas[1] ; fz[21] double mutants show a novel phenotype, with bristles on the notum oriented towards the anterior. (Olguín et al., 2011) Heterozygous Rab23[51] significantly increases the number of multiple wing hairs in fz[21] mutants. The double mutant combination fz[21], Rab23[51] displays a very high number of multiple wing hairs and strong hair orientation defects. (Pataki et al., 2010) Vang[stbm-6] clones in the pupal wing in a uniformly fz[21] background show no difference in the time of prehair initiation between single and double mutant tissue. fz[21] clones in the pupal wing in a uniformly Vang[stbm-6] background show prehairs initiating sooner in Vang[stbm-6] mutant tissue than in Vang[stbm-6] ; fz[21] double mutant tissue. (Strutt and Warrington, 2008) The planar cell polarity defects seen in Egfr[f11]/Egfr[t1] eyes are partially suppressed by fz[21]/+. The symmetrical ommatidium phenotype seen in fz[19]/fz[21] flies is suppressed by Egfr[top-18A]/+. The frequency of rotation defects in the ommatidia of Egfr[f24]/+ flies is reduced to 12% or 18% respectively if they are also carrying fz[21]/+. (Weber et al., 2008) As is seen with fz[21] clones alone, fz[21] Vang[stbm-6] double mutant clones non-autonomously reorient the wing hairs of neighbouring wild type cells so that they point towards the clone. Expression of fz::fz2[2-1-1.tub.T:Hsap\MYC] fails to rescue the non-autonomous effects of fz[23]/fz[21] mutant clones on the orientation of the wing hairs of neighbouring wild type cells. (Wu and Mlodzik, 2008) fz[21] fz2[C1] double homozygous clones in the dorsal air sac primordium grow normally and populate the tip of the air sac primordium to the same degree as wild-type clones. (Cabernard and Affolter, 2005) No evidence of a domineering nonautonomy wing hair polarity phenotype is seen in fz²¹ clones in a frtz^unspecified background. (Collier et al., 2005) fz2^{ΔCRD.αTub84B.T:wg,T:Ivir\HA1} suppresses loss of naked cuticle in fz²¹/fz²¹, Df(3L)fz2/+ embryos. The additional presence of fz3^G10/fz3^G10 has no effect on this phenotype or its suppression. Somatic clones of cells homozygous for fz²¹ and fz2^C2 at the anterior wing margin block the formation of wing margin bristles and cause wing notching. These phenotypes are fully suppressed by fz^{1-1-1.αTub84B}, fz2^αTub84B.PC, fz^{ΔCRD.αTub84B.T:wg,T:Ivir\HA1} or fz2^{ΔCRD.αTub84B.T:wg,T:Ivir\HA1}. (Chen et al., 2004) The planar polarity phenotype in the abdomen of fz²¹/fz²¹ flies is largely unaffected when the whole animal is also pk^{sple.Scer\UAS} and Scer\GAL4^hh-Gal4 or pk^pk-sple-13/pk^pk-sple-13. The loss of polarity seen adjacent to Vang^stbm-6 somatic clones in the adult abdomen is suppressed by fz²¹/fz²¹. The zone of reversed polarity at the back and behind the anterior compartment of each abdominal segment of stan^Scer\UAS.cUa; Scer\GAL4^ptc-559.1 adults is suppressed by fz³⁷/fz²¹. (Lawrence et al., 2004) When a wing clone is doubly mutant for fz²¹ and in^IH56, domineering nonautonomy of polarity defect of fz²¹ is unaffected. When a wing clone is doubly mutant for fz²¹ and in^IH56 in a genetic background deficient for in, domineering nonautonomy of polarity defect of fz²¹ is suppressed, though polarity defect within the clone is not. (Lee and Adler, 2002) Double mutants with Df(3L)fz2 show variable segmentation defects ranging from a few extra denticles in the posterior part of some segments to complete replacement of naked cuticle with denticles. This phenotype is reminiscent of that for wg loss of function. The expression of arm^ΔN.Scer\UAS driven by Scer\GAL4^da.G32 completely overrides this phenotype, causing naked cuticle. Double mutants with Df(3L)fz2 show a complete loss of RP2 neurons in all hemisegments. In double mutants of fz²¹ or fz²³ with Df(3L)fz2 cardiac precursors are missing. Double mutants of fz²¹ or fz²³ with Df(3L)fz2 lack the normal number of midgut constrictions. Overexpression of wg^Scer\UAS.cLb by Scer\GAL4^da.G32 has no effect on the cuticle phenotype of double mutants of fz²¹ or fz²³ with Df(3L)fz2. (Bhanot et al., 1999)
Xenogenetic Interactions
Statement Reference
Complementation & Rescue Data
Rescued by	fz²¹ is rescued by Scer\GAL4^da.G32/fz^Scer\UAS.cAa (Bhanot et al., 1999) fz²³/fz²¹ is rescued by fz^1-1-1.tub.W (Wu et al., 2004) fz²³/fz²¹ is rescued by fz^{565Stop.tub.T:Hsap\MYC} (Wu et al., 2008) fz²³/fz²¹ is rescued by fz^{tub.T:Hsap\MYC} (Wu et al., 2008)
Not rescued by	fz²³/fz²¹ is not rescued by fz^{M469R.tub.T:Hsap\MYC} (Wu et al., 2008) fz²³/fz²¹ is not rescued by fz^{ΔCRD.tub.T:Hsap\MYC} (Wu and Mlodzik, 2008) fz²³/fz²¹ is not rescued by fz^{ΔSKT.tub.T:Hsap\MYC} (Wu et al., 2008) fz²³/fz²¹ is not rescued by fz^{ΔSWRNF.tub.T:Hsap\MYC} (Wu et al., 2008)
Comments	Expression of fz[ΔCRD.tub.T:Hsap\MYC] fails to rescue the non-autonomous effects of fz[23]/fz[21] mutant clones on the orientation of the wing hairs of neighbouring wild type cells. 92.3% of clones have surrounding wild type cells pointing towards the clone. (Wu and Mlodzik, 2008) Expression of fz[tub.T:Hsap\MYC] rescues the wing hair phenotype seen in fz[21]/fz[23] mutants. Expression of fz[ΔSWRNF.tub.T:Hsap\MYC] fails to rescue the wing hair phenotype seen in fz[21]/fz[23] mutants. Expression of fz[ΔSKT.tub.T:Hsap\MYC] fails to rescue the wing hair phenotype seen in fz[21]/fz[23] mutants. Expression of fz[M469R.tub.T:Hsap\MYC] fails to rescue the wing hair phenotype seen in fz[21]/fz[23] mutants. Expression of fz[565Stop.tub.T:Hsap\MYC] rescues the wing hair phenotype seen in fz[21]/fz[23] mutants. (Wu et al., 2008)
Stocks ( 1 )
Bloomington	41787 fz²¹ th¹ st¹/TM3, Sb¹
Notes on Origin
Discoverer	Adler.

External Crossreferences & Linkouts
Other Crossreferences

Linkouts

Synonyms & Secondary IDs ( 6 )
Reported As
Symbol Synonym	21 (Weber et al., 2012) fz²¹ (Lawrence et al., 2004, Classen et al., 2005, Casal et al., 2006, Strutt and Warrington, 2008, Pataki et al., 2010, Bastock and Strutt, 2007, Strutt and Strutt, 2008) fz^- (Casal et al., 2006) fz^P21 (Piddini et al., 2005, Povelones et al., 2005, Collier et al., 2005, Han et al., 2005, Katanaev et al., 2005, Chen et al., 2004, Wu et al., 2004, Lee and Adler, 2004, Lee and Adler, 2002, Cohen et al., 2002, Gao et al., 2000, Adler et al., 2000, Bhanot et al., 1999, Jones et al., 1996, Walters et al., 2006, Hermle et al., 2013, Wu et al., 2008, Weber et al., 2008, Chung et al., 2009, Doyle et al., 2008, Blanco et al., 2009, Djiane and Mlodzik, 2010, Lu et al., 2010, Cho and Fischer, 2011, Wu and Mlodzik, 2008) fz^p21 (Olguín et al., 2011, Cabernard and Affolter, 2005, Olguín et al., 2011, Gault et al., 2012) P21 (Weber et al., 2012)
Name Synonym
Secondary FlyBase IDs

References ( 38 )

Generate a list of	All references relating to this allele ( 38 )

List References by type	Research paper ( 37 ) Supplementary material ( 1 )
Recent research papers ( 5 )
	Hermle et al., 2013, EMBO J. 32(2): 245--259 Drosophila ATP6AP2/VhaPRR functions both as a novel planar cell polarity core protein and a regulator of endosomal trafficking. [FBrf0220613] Gault et al., 2012, J. Cell Biol. 196(5): 605--621 Drosophila CK1-γ, gilgamesh, controls PCP-mediated morphogenesis through regulation of vesicle trafficking. [FBrf0217630] Weber et al., 2012, Genetics 191(1): 145--162 Novel regulators of planar cell polarity: a genetic analysis in Drosophila. [FBrf0218210] Cho and Fischer, 2011, Development 138(7): 1349--1359 Ral GTPase promotes asymmetric Notch activation in the Drosophila eye in response to Frizzled/PCP signaling by repressing ligand-independent receptor activation. [FBrf0213208] Olguín et al., 2011, Curr. Biol. 21(3): 236--242 Intertissue mechanical stress affects frizzled-mediated planar cell polarity in the Drosophila notum epidermis. [FBrf0212947] Show all research papers ...

Allele Dmel\fz21

Allele Dmel\fz²¹