Homozygous clones in the retina result in a mildly disorganised ommatidial pattern and a slight reduction in eye size. Females carrying homozygous germ line clones are completely sterile and fail to lay eggs. Most egg chambers in these females can proceed normally through the early and middle stages of oogenesis, but arrest at the stage when the nurse cells rapidly transfer their contents to the oocyte. The arrested egg chambers have enlarged nurse cells and a small poorly developed oocyte. The actin filaments that normally tether the nurse cell nuclei mostly fail to form. The nurse cell nuclei are inappropriately positioned close to the ring canals and are sometimes seen to stretch partially through the ring canals.