At 21oC, wings are curved downward and held out slightly, they may be crumpled or not quite unfolded. Tarsi are weak but usually not deformed. Expression is more extreme at 27oC and overlaps wild-type at 17oC. Viability is very low (near-lethal) at 27oC and is fair at 17oC. rk1/rk2 is intermediate between rk1/rk1 and rk2/rk2, with the wings arched and papery, sometimes not unfolded.