Transheterozygotes show no lethality throughout normal larval development. The larvae continue as third instar larvae for several days longer than their heterozygous siblings, eventually dying as third instar larvae or (in some allelic combinations) as early pupae. Third instar larvae have reduced optic lobes in the larval brain and reduced or missing imaginal discs for most allelic combinations.
Strong enhancer of variegation at the lt locus. Also enhances variegation at the l(2)40Fa and l(2)40Ff loci.