686bp deletion resulting from the imprecise excision of a P element that is reported to extend from -715bp to -30bp relative to the wdn transcription start site. 3R:28811338 was used as the transcription start site (FBrf0048671).
Homozygous flies have a number of defects; young flies generally hold their wings at approximately 45o above horizontal, whereas older flies (and occasionally young flies) have drooping wings that prevent the fly from walking normally. The anterior cross vein is often either deformed or eliminated by increased proximity of veins L3 and L4. The posterior cross vein is often interrupted, with the anterior part is missing. Some of the macrochaetae, most frequently the scutellar, postalar and dorsocentral, may be duplicated or triplicated, with 2-6 extra bristles per fly. This extra bristle phenotype is stronger in females than males. Homozygous females give no viable progeny, whatever the genotype of the male they are mated to. Approximately 75% of the eggs laid by these females appear to be unfertilised and undergo no nuclear division. Approximately 25% undergo nuclear cleavages and cellularise. Morphological movements leading to gastrulation and germ-band extension are severely disrupted. In all cases these animals die as embryos. 1-3% of the embryos do not show developmental alterations before or during gastrulation and germ-band extension, but die as morphologically normal prehatching embryos or first or second instar larvae.