sogYL26/+; tokΔ2-41/tokE1 escapers exhibit partial posterior crossveins with significantly more crossvein material than tokΔ2-41/tokE1 escapers. Further, crossveins are almost completely restored in sogYL26/sogP129D; tokΔ2-41/tokE1 animals.
Only 39% of sogP129D/+ embryos derived from MadES1/+ females survive to adulthood. sogP129D embryos derived from MadES1/+ females have internalised filzkorper and lack head elements, indicative of weak ventralisation. The amnioserosa is reduced. sogP129D ; zen1 double mutant embryos die with a partially ventralised phenotype at 25oC.