The cuticle occasionally has a dorsal hole in homozygous embryos. There are often fewer pericardial cells than in wild-type embryos, resulting in distinct gaps in the heart tube. The pericardial cells appear to dissociate randomly and the heart tube often appears to curve off towards the lateral side of the embryo.
Lethality occurs during third instar larval or pupal stages. Mutants do not show any visible disc abnormalities.