Homozygotes show significant lethality during all stages examined (embryogenesis to pharate adult). Some homozygous adults are able to eclose, but they die 2 to 3 days later. Homozygous pharate adults have malformed legs which may fail to extend to the tip of the abdomen but remain organised. The femur may contain constrictions. 91% of mutant mid-prepupae have at least one malformed leg, 94% of mutant pupae and adults have at least one malformed leg.