Lethality occurs during third instar larval or pupal stages. Mutants exhibit visible disc abnormalities. In tergites, clones with normal size and frequency are recovered. Clones cannot be recovered in the wing disc.
Brain, imaginal disc and blood cell neoplastic phenotype.
Brain is of normal size but abnormal shape. Wing and haltere discs rarely showing hyperplastic outgrowths, folding abnormal. Lymph gland retains lobular structure.