Adult expression is observed in a cluster of five or six tangential large field neurons with cell bodies located in the dorso-caudal cellular cortex lateral to the calyx of the mushroom body. The axons of these neurons arborize in the rostro-dorsal protocerebrum, near the tip of the α'-lobe of the mushroom body, and dorsally in the fan-shaped body. Additional expression is observed in the lamina of the optic lobe, in the ocelli, and in the antennal nerve.