Homozygotes die during the 2nd to 3rd larval instar without any discernible morphological defects. Homozygous egg chambers in female germ-line clones do not complete oogenesis. Most of the egg chambers degenerate during mid-oogenesis, such that the cell membranes of germline cells are collapsed and ring canals and nurse cell nuclei are concentrated to form large aggregates within egg chambers. Nurse cell nuclei in these egg chambers are then fragmented into small pieces. Occasionally, egg chambers which have progressed to stage 10 are seen. In these egg chambers, the oocyte does not grow normally the and the nurse cell nuclei are displaced posteriorly relative to the anterior border of columnar follicle cells (which normally surround the oocyte at stage 10). The egg chambers are morphologically normal until stage 4-5 and the oocyte fails to grow normally from stage 6 onwards, at which stage the egg chambers begin to degenerate.

me31B^Δ1/me31B[+] is an enhancer of dorsal appendage | maternal effect phenotype of BicC⁴