About 50% of Grip84^PG36 hemizygous males reach the adult stage after a 2 day delay compared to controls. The resulting adults are sterile and have abnormalities in the abdominal cuticle and pattern of thoracic macrochaetae. Most male germline cysts in these animals have only 16 cells, suggesting that the two of the rounds divisions have not occurred properly. Onion stage spermatids from these males frequently have multiple nuclei and/or multiple Nebenkern. Nebenkern and nuclei are often abnormally sized. In primary spermatocytes from these animals, microtubule asters are not visible and most spindles are highly abnormal: some are devoid of any polarity, others are monopolar with microtubules diverging to form an umbrella shape or converging to form a cone. Marker analysis suggests defects in pericentriolar matrix formation.

In Grip84^PG36 homozygous late third instar larvae, the mitotic index of larval brain neuroblasts is elevated is elevated 4-5 fold over wild-type levels: cells accumulate in pro-metaphase and metaphase; fewer cells are found in anaphase or telophase. Many of these neuroblasts have overcondensed chromosomes and disorganised metaphase plates.