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Gene Dmel\osp

General Information
Symbol	Dmel\osp	Species	D. melanogaster
Name	outspread	Annotation symbol	CG3479
Feature type	protein_coding_gene	FlyBase ID	FBgn0003016
Gene Model Status	Current	Stock availability	28 publicly available
Also Known As	BG:DS01486.7
Genomic Location
Chromosome (arm)	2L	Recombination map	2-50.1
Cytogenetic map	35B3-35B4	Sequence location	2L:14,599,776..14,689,325 [-]
Genomic Maps Select View:Gene Models/EvidenceExpression/RegulationMegaViewGene Disruptions, Stocks and Reagents Beta	Decorated FastA GenBankGFF Gene region Extended Gene region CDSIntronsExonsTranslationsTranscripts3' UTR5' UTR
Summary Information
Automatically generated summary See sections below for more information	The gene outspread is referred to in FlyBase by the symbol Dmel\osp (CG3479, FBgn0003016). It is a protein_coding_gene from Drosophila melanogaster. An electronic pipeline based on InterPro domains suggests that it has the molecular function: phospholipid binding. The biological processes in which it is involved are not known. 59 alleles are reported. The phenotypes of these alleles are annotated with: wing; thorax; crystal cell. It has 4 annotated transcripts and 4 annotated polypeptides. Protein features are: Pleckstrin homology domain; Pleckstrin homology-like domain. Summary of modENCODE Temporal Expression Profile: Temporal profile ranges from a peak of high expression to a trough of low expression. Peak expression observed during late pupal stages. Summary of FlyAtlas Anatomical Expression Data: Expression at moderate levels in the following post-embryonic organs or tissues: adult head, adult eye, larval/adult central nervous system, larval hindgut, larval Malpighian tubules, larval/adult fat body, larval trachea, adult female reproductive system, larval/adult carcass. Comments on Affy2 ProbeSet: ProbeSet 1627054_at completely aligns to an exonic region common to each of the 4 FlyBase-annotated transcript isoforms of osp. Gene sequence location is 2L:14599776..14689325.
User Contributed Data
	Beta version, feedback welcome
Phenotypic Description from the Red Book (Lindsley & Zimm 1992)
Gene/Allele symbols may differ from current usage	*alw: arclike wing Wings evenly bent downward at tips. Overlaps wild type. RK2. osp: outspread (M. Ashburner) Wings held out at 45 to body axis. Strong alleles, and homozygous deficiencies show, in addition, a "tenting" of the wings so that they resemble those of arc. Df(2L)osp homozygous viable and fertile, though viability reduced in crowded cultures.
Recent Updates
Description	What does this section display? This section contains items that were added to this record for each release. It currently only tracks new links between this FlyBase report and other FlyBase data classes (e.g. genes, references, stocks) or controlled vocabulary terms (e.g. GO, anatomy terms). What does this section not display? This section does not currently display links that were removed or gene model changes.
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FB2013_03
FB2013_02
All updates	Click here to see a list of all updates to this record from FB2010_08 and on.
Detailed Mapping Data
FlyBase Computed Cytological Location
Cytogenetic map Evidence for location 35B3-35B4   Limits computationally determined from genome sequence between P{EP}elBEP2039&P{PZ}osprJ571 and P{lacW}Su(H)k07904
Experimentally Determined Cytological Location
Cytogenetic map Notes References 35B1-35B4   (determined by in situ hybridisation) 35B3--5 (determined by in situ hybridisation)   (Ashburner et al., 1999) 35B1-35B4   (determined by in situ hybridisation)   (Spradling et al., 1999) 35B1-35B4     (BDGP Project Members, 1994-1999)
Experimentally Determined Recombination Data
Location	2-50.1
Left of (cM)	Adh (Lindsley and Zimm, 1992) rd (Woodruff and Ashburner, 1979) Bl (Sturtevant, 1948)
Right of (cM)	S (Sturtevant, 1948) Adh (Woodruff and Ashburner, 1979)
Notes	Maps very near b. (Sturtevant, 1948)
Gene Model & Products
Please see the GBrowse view of Dmel\osp for information on other features To submit a correction to a gene model please use the Contact FlyBase form
Comments on Gene Model
	DGC clone LD14119 appears problematic: incomplete CDS
Transcript Data
Annotated Transcripts
Name FlyBase ID RefSeq ID Length (nt) Associated CDS (aa) osp-RC FBtr0114465  NM_001144354   5902   1386 osp-RD FBtr0301494  NM_165089   6224   1557 osp-RE FBtr0301495  NM_078843   5875   1377 osp-RF FBtr0301496  NM_001169504   6251   1566
Additional Transcript Data & Comments
Reported size (kB)
Comments
External Data
Crossreferences
Polypeptide Data
Annotated Polypeptides
Name FlyBase ID Predicted MW (kDa) Length (aa) Theoretical pI RefSeq ID GenBank protein osp-PC   FBpp0112957   156.2   1386   6.63     NP_001137826   ACL83032 osp-PD   FBpp0290709   174.5   1557   6.58     NP_723879   AAF53402 osp-PE   FBpp0290710   155.2   1377   6.63     NP_523567   AAN10878 osp-PF   FBpp0290711   175.5   1566   6.58     NP_001162975   ACZ94261
Additional Polypeptide Data & Comments
Reported size (kDa)
Comments
External Data
Linkouts
Crossreferences	InterPro domains - A database of protein families, domains, and functional sites • Pleckstrin homology domain (IPR001849) Pleckstrin homology-like domain (IPR011993)
Sequences Consistent with the Gene Model
DDBJ / EMBL / GenBank	DNA sequence Protein sequence Name AA140663   AA140932   AA141121   AA141124   AA141127   AA141143   AA141196   AA141199   AA141201   AA141234   AA141323   AA141354   AA141371   AA141408   AA141422   AA141464   AA141478   AA141489   AA141497   AA141560   AA141562   AA141607   AA141609   AA141611   AA141676   AA141689   AA141707   AA141711   AA141735   AA141736   AA141743   AA141801   AA141834   AA141849   AA141881   AA142178   AA439264   AA439528   AA440163   AA441103   AA736187   AE014134 AAF53402   AE014134 AAN10878   AE014134 ACL83032   AE014134 ACZ94261   AE003410 AAF44880   AE003410 AAF44881   AI124308   AI296338   AI387761   AI531290   AI546631   AQ026050   AQ026134   AQ073307   AW942302   AX664501   AY118512 AAM49881   BG640306   BI142244   BI165760   BI170191   BI352942   BI364495   BI564851   BI565487   BI571886   BI574131   BI575214   BI576026   BI577030   BI582454   BI583280   BI583851   BI583993   BI586861   BI588406   BI588459   BI588663   BI588766   BI588884   BI589090   BI589239   BI589243   BI590672   BI590731   BI592794   BI593032   BI604445   BI605511   BI606547   BI607294   BI607877   BI607977   BI608784   BI608982   BI610522   BI610661   BI611018   BI611097   BI611159   BI611230   BI611710   BI612977   BI613020   BI613394   BI613754   BI615370   BI615509   BI616145   BI616581   BI616610   BI616906   BI617028   BI617090   BI617439   BI617684   BI618824   BI619653   BI620053   BI620538   BI620781   BI621147   BI621284   BI622500   BI622740   BI623088   BI623093   BI623319   BI623627   BI624745   BI625317   BI625766   BI625785   BI625802   BI626350   BI626426   BI626450   BI626871   BI627441   BI627518   BI628367   BI628419   BI628426   BI628436   BI628544   BI629001   BI629190   BI629211   BI629381   BI630149   BI630924   BI631188   BI631763   BI631785   BI631848   BI631995   BI634197   BI634339   BI634950   BI635165   BI636622   BI641728   BP546474   BT011069   BT030131   BT031256   CA804042   CK131876   CK135090   CK660751   CO153056   CO153494   CO177555   CO179042   CO179169   CO179305   CO180091   CO186002   CO187196   CO187484   CO187678   CO187710   CO187913   CO189129   CO190350   CO190469   CO190773   CO190982   CO195863   CO261955   CO262568   CO263072   CO263244   CO263326   CO263592   CO264634   CO266208   CO266621   CO267454   CO267553   CO268027   CO268650   CO269696   CO271850   CO271971   CO273023   CO274206   CO274786   CO274948   CO275125   CO275338   CO275570   CO275968   CO276551   CO277967   CO278383   CO278479   CO278943   CO279082   CO280443   CO281966   CO282253   CO282275   CO282855   CO283735   CO284008   CO284265   CO284404   CO284486   CO286623   CO286760   CO286927   CO290009   CO290180   CO290679   CO290759   CO291316   CO291587   CO291788   CO291860   CO292855   CO293512   CO293878   CO294202   CO294380   CO294540   CO295238   CO295444   CO295750   CO297010   CO297570   CO298241   CO300381   CO300478   CO301305   CO302049   CO302309   CO302349   CO302601   CO302681   CO303318   CO303356   CO303980   CO304026   CO304946   CO307131   CO308453   CO309390   CO309547   CO309805   CO310162   CO310496   CO311647   CO311693   CO311933   CO312111   CO312182   CO314165   CO315577   CO315650   CO316352   CO316667   CO317096   CO317213   CO317312   CO317355   CO319487   CO319523   CO320976   CO320977   CO321140   CO322033   CO323072   CO324635   CO324658   CO324776   CO326836   CO326862   CO327048   CO327604   CO328187   CO328257   CO328314   CO328871   CO329111   CO330975   CO331792   CO332493   CO332733   CO332776   CO337673   CO338920   CO339169   CO339604   CO340021   CO340197   CO340225   CO340355   CO340387   CO340748   CO340851   CO340882   CO340913   CO340947   CO341122   CO341648   CO341778   CO341926   CO341986   CO342406   CO342752   CO343038   CO343368   CO343580   CO343721   CO344264   CO344379   CO344766   CO344944   CO344971   CZ466231   CZ469498   CZ470194   CZ475345   CZ477844   CZ480752   CZ480753   CZ480755   CZ485760   CZ485761   CZ488084   CZ488085   CZ488200   CZ488201   CZ489874   EC061058   EC062076   EC062387   EC065381   EC071467   EC072200   EC074826   EC075542   EC076005   EC080166   EC086115   EC201928   EC207005   EC208007   EC208228   EC208671   EC211693   EC213663   EC219558   EC219559   EC219626   EC228236   EC229563   EC237346   EC241902   EC242632   EC246124   EC247570   EC249935   EC253499   EC256474   EC258434   EC260525   EV576899   EV580573   EV594450   EV599365   GH942097   GH943761   GH943919   GH944707   GH945150   GH945765   GH946168   GH946479   GH947141   GH948871   GH951818   GH952169   GH952205   GH952318   GH954267   GH954634   GH955297   GH955660   GH956562   GH959863   GH959896   GH960098   GH960879   GH961094   GH961887   GH962968   GH963415   GH963630   GH966449   GH966478   GH966911   GH967243   GH967887   GH968786   GH969283   GH969683   GH969832   GH970229   GH970376   GH972799   GH974765   GH975033   GH977259   GH978697   GH979055   GH981064   GH983105   GH983112   GH983459   GH983968   GH984319   GH984640   GH984744   GH984846   GH985051   M17837 AAA70211   M19547 AAA70209   Z00030 CAA77329
UniProtKB/Swiss-Prot	Q27421
UniProtKB/TrEMBL
Mapped Features
Mapped Features have been reorganized, please see this article for details. Additional mapped features and mutations can be found on GBrowse or related reports. Type Symbol & Location Additional Notes References
External Data
Linkouts
Crossreferences
Expression Data
Transcript Expression
in situ Stage Tissue/Position (including subcellular localization) Reference embryonic stage 11 -- 12   salivary gland body primordium   (Fisher et al., 2012) embryonic stage 13 -- 16   embryonic foregut   (Fisher et al., 2012) embryonic hypopharynx   (Fisher et al., 2012) embryonic/larval salivary gland   (Fisher et al., 2012) embryonic/larval salivary gland body   (Fisher et al., 2012)
Additional Descriptive Data
Marker for
Subcellular Localization
CV Term
Polypeptide Expression
mass spectroscopy Stage Tissue/Position (including subcellular localization) Reference day 5 of adulthood & male -- day 7 of adulthood & male   spermatozoon   (Wasbrough et al., 2010)
Additional Descriptive Data
Marker for
Subcellular Localization (GO Cellular Component)
CV term Evidence References
Expression Deduced from Reporters
High-Throughput Expression Data
Associated Tools
Reference	See Gelbart and Emmert, 2010.10.13 for analysis details and data files for all genes.
FlyAtlas Anatomy Microarray
FlyAtlas Anatomical Expression Data (FlyAtlas-RNA.adult) (FlyAtlas-RNA.larva)    Styles Linear Logarithmic Heatmap Back-to-back    Scales gene max expression Moderate expression bin max High level expression bin max Very high expression bin max Summary of FlyAtlas Anatomical Expression Data: Expression at moderate levels in the following post-embryonic organs or tissues: adult head, adult eye, larval/adult central nervous system, larval hindgut, larval Malpighian tubules, larval/adult fat body, larval trachea, adult female reproductive system, larval/adult carcass. [download data (TSV)] Guide to FlyAtlas expression level colors   No expression (0 - 9.999)   Low expression (10 - 99.999)   Moderate expression (100 - 499.999)   High level expression (500 - 999.999)   Very high expression (>999.999) Linear, scaled to maximum expression level Tissue   Expression Level Larval Central Nervous System    120.925 Larval Midgut    73.4 Larval Hindgut    224.8 Larval Malpighian Tubules    147.5 Larval Fat Body    161.7 Larval Salivary Gland    69.4 Larval Trachea    143.4 Larval Carcass    106.975 Adult Head    149.3 Adult Eye    180.225 Adult Brain    118.4 Adult Thoracic-Abdominal Ganglion    192.6 Adult Crop    78.7 Adult Midgut    33.5 Adult Hindgut    97.3 Adult Malpighian Tubules    82.3 Adult Fat Body    112.9 Adult Salivary Gland    80.5 Adult Heart    60.775 Adult VirginFemale Spermatheca    223.6 Adult InseminatedFemale Spermatheca    215 Adult Ovary    276.3 Adult Testis    56.6 Adult Male Accessory Gland    60 Adult Carcass    107.2 Expression Level Scale  None   Low   Moderate  Linear, scaled to Moderate expression Tissue   Expression Level Larval Central Nervous System    120.925 Larval Midgut    73.4 Larval Hindgut    224.8 Larval Malpighian Tubules    147.5 Larval Fat Body    161.7 Larval Salivary Gland    69.4 Larval Trachea    143.4 Larval Carcass    106.975 Adult Head    149.3 Adult Eye    180.225 Adult Brain    118.4 Adult Thoracic-Abdominal Ganglion    192.6 Adult Crop    78.7 Adult Midgut    33.5 Adult Hindgut    97.3 Adult Malpighian Tubules    82.3 Adult Fat Body    112.9 Adult Salivary Gland    80.5 Adult Heart    60.775 Adult VirginFemale Spermatheca    223.6 Adult InseminatedFemale Spermatheca    215 Adult Ovary    276.3 Adult Testis    56.6 Adult Male Accessory Gland    60 Adult Carcass    107.2 Expression Level Scale  None   Low   Moderate   High  Linear, scaled to High level expression Tissue   Expression Level Larval Central Nervous System    120.925 Larval Midgut    73.4 Larval Hindgut    224.8 Larval Malpighian Tubules    147.5 Larval Fat Body    161.7 Larval Salivary Gland    69.4 Larval Trachea    143.4 Larval Carcass    106.975 Adult Head    149.3 Adult Eye    180.225 Adult Brain    118.4 Adult Thoracic-Abdominal Ganglion    192.6 Adult Crop    78.7 Adult Midgut    33.5 Adult Hindgut    97.3 Adult Malpighian Tubules    82.3 Adult Fat Body    112.9 Adult Salivary Gland    80.5 Adult Heart    60.775 Adult VirginFemale Spermatheca    223.6 Adult InseminatedFemale Spermatheca    215 Adult Ovary    276.3 Adult Testis    56.6 Adult Male Accessory Gland    60 Adult Carcass    107.2 Expression Level Scale  None   Low   Moderate   High   Very high  Linear, scaled to Very high expression Tissue   Expression Level Larval Central Nervous System    120.925 Larval Midgut    73.4 Larval Hindgut    224.8 Larval Malpighian Tubules    147.5 Larval Fat Body    161.7 Larval Salivary Gland    69.4 Larval Trachea    143.4 Larval Carcass    106.975 Adult Head    149.3 Adult Eye    180.225 Adult Brain    118.4 Adult Thoracic-Abdominal Ganglion    192.6 Adult Crop    78.7 Adult Midgut    33.5 Adult Hindgut    97.3 Adult Malpighian Tubules    82.3 Adult Fat Body    112.9 Adult Salivary Gland    80.5 Adult Heart    60.775 Adult VirginFemale Spermatheca    223.6 Adult InseminatedFemale Spermatheca    215 Adult Ovary    276.3 Adult Testis    56.6 Adult Male Accessory Gland    60 Adult Carcass    107.2 Expression Level Scale  Very high  log, scaled to maximum expression level Tissue   Expression Level Larval Central Nervous System    120.925 Larval Midgut    73.4 Larval Hindgut    224.8 Larval Malpighian Tubules    147.5 Larval Fat Body    161.7 Larval Salivary Gland    69.4 Larval Trachea    143.4 Larval Carcass    106.975 Adult Head    149.3 Adult Eye    180.225 Adult Brain    118.4 Adult Thoracic-Abdominal Ganglion    192.6 Adult Crop    78.7 Adult Midgut    33.5 Adult Hindgut    97.3 Adult Malpighian Tubules    82.3 Adult Fat Body    112.9 Adult Salivary Gland    80.5 Adult Heart    60.775 Adult VirginFemale Spermatheca    223.6 Adult InseminatedFemale Spermatheca    215 Adult Ovary    276.3 Adult Testis    56.6 Adult Male Accessory Gland    60 Adult Carcass    107.2 Expression Level Scale  None   Low   Moderate  log, scaled to Moderate expression Tissue   Expression Level Larval Central Nervous System    120.925 Larval Midgut    73.4 Larval Hindgut    224.8 Larval Malpighian Tubules    147.5 Larval Fat Body    161.7 Larval Salivary Gland    69.4 Larval Trachea    143.4 Larval Carcass    106.975 Adult Head    149.3 Adult Eye    180.225 Adult Brain    118.4 Adult Thoracic-Abdominal Ganglion    192.6 Adult Crop    78.7 Adult Midgut    33.5 Adult Hindgut    97.3 Adult Malpighian Tubules    82.3 Adult Fat Body    112.9 Adult Salivary Gland    80.5 Adult Heart    60.775 Adult VirginFemale Spermatheca    223.6 Adult InseminatedFemale Spermatheca    215 Adult Ovary    276.3 Adult Testis    56.6 Adult Male Accessory Gland    60 Adult Carcass    107.2 Expression Level Scale  None   Low   Moderate   High  log, scaled to High level expression Tissue   Expression Level Larval Central Nervous System    120.925 Larval Midgut    73.4 Larval Hindgut    224.8 Larval Malpighian Tubules    147.5 Larval Fat Body    161.7 Larval Salivary Gland    69.4 Larval Trachea    143.4 Larval Carcass    106.975 Adult Head    149.3 Adult Eye    180.225 Adult Brain    118.4 Adult Thoracic-Abdominal Ganglion    192.6 Adult Crop    78.7 Adult Midgut    33.5 Adult Hindgut    97.3 Adult Malpighian Tubules    82.3 Adult Fat Body    112.9 Adult Salivary Gland    80.5 Adult Heart    60.775 Adult VirginFemale Spermatheca    223.6 Adult InseminatedFemale Spermatheca    215 Adult Ovary    276.3 Adult Testis    56.6 Adult Male Accessory Gland    60 Adult Carcass    107.2 Expression Level Scale  None   Low   Moderate   High   Very high  log, scaled to Very high expression Tissue   Expression Level Larval Central Nervous System    120.925 Larval Midgut    73.4 Larval Hindgut    224.8 Larval Malpighian Tubules    147.5 Larval Fat Body    161.7 Larval Salivary Gland    69.4 Larval Trachea    143.4 Larval Carcass    106.975 Adult Head    149.3 Adult Eye    180.225 Adult Brain    118.4 Adult Thoracic-Abdominal Ganglion    192.6 Adult Crop    78.7 Adult Midgut    33.5 Adult Hindgut    97.3 Adult Malpighian Tubules    82.3 Adult Fat Body    112.9 Adult Salivary Gland    80.5 Adult Heart    60.775 Adult VirginFemale Spermatheca    223.6 Adult InseminatedFemale Spermatheca    215 Adult Ovary    276.3 Adult Testis    56.6 Adult Male Accessory Gland    60 Adult Carcass    107.2 Expression Level Scale  None   Low   Moderate   High   Very high  Heatmap Tissue   Expression Level Larval Central Nervous System     Larval Midgut     Larval Hindgut     Larval Malpighian Tubules     Larval Fat Body     Larval Salivary Gland     Larval Trachea     Larval Carcass     Adult Head     Adult Eye     Adult Brain     Adult Thoracic-Abdominal Ganglion     Adult Crop     Adult Midgut     Adult Hindgut     Adult Malpighian Tubules     Adult Fat Body     Adult Salivary Gland     Adult Heart     Adult VirginFemale Spermatheca     Adult InseminatedFemale Spermatheca     Adult Ovary     Adult Testis     Adult Male Accessory Gland     Adult Carcass     FlyAtlas Organ/Tissue Expression, larval vs. adult Larval Expression Level Tissue Adult Expression Level   NA  Head    149.3   NA  Eye    180.225   NA  Brain    118.4   120.925  Central Nervous System    NA   NA  Thoracic-Abdominal Ganglion    192.6   NA  Crop    78.7   73.4  Midgut    33.5   224.8  Hindgut    97.3   147.5  Malpighian Tubules    82.3   161.7  Fat Body    112.9   69.4  Salivary Gland    80.5   NA  Heart    60.775   143.4  Trachea    NA   NA  VirginFemale Spermatheca    223.6   NA  InseminatedFemale Spermatheca    215   NA  Ovary    276.3   NA  Testis    56.6   NA  Male Accessory Gland    60   106.975  Carcass    107.2 FlyAtlas Anatomical Expression Data (Chintapalli et al., 2007)
modENCODE Anatomy RNA-Seq
modENCODE Tissue Expression Data (modENCODE_mRNA-Seq_tissues)    Styles Linear Logarithmic Heatmap    Scales gene max expression Moderate expression bin max High expression bin max Extremely high expression bin max [download data (TSV)] Guide to modENCODE expression level colors   No/Extremely low expression (0 - 0)   Very low expression (1 - 3)   Low expression (4 - 10)   Moderate expression (11 - 25)   Moderately high expression (26 - 50)   High expression (51 - 100)   Very high expression (101 - 1000)   Extremely high expression (>1000) Linear, scaled to maximum expression level Tissue   Expression Level imaginal disc, larvae L3 wandering    11 central nervous system, larvae L3    15 central nervous system, pupae P8    37 head, virgin 1-day female    16 head, virgin 4-day female    16 head, virgin 20-day female    8 head, mated 1-day female    17 head, mated 4-day female    12 head, mated 20-day female    10 head, mated 1-day male    21 head, mated 4-day male    20 head, mated 20-day male    17 salivary gland, larvae L3 wandering    6 salivary gland, white prepupae    3 digestive system, larvae L3 wandering    6 digestive system, 1-day adult    3 digestive system, 4-day adult    5 digestive system, 20-day adult    3 fat body, larvae L3 wandering    9 fat body, white prepupae    13 fat body, pupae P8    34 carcass, larvae L3 wandering    15 carcass, 1-day adult    14 carcass, 4-day adult    14 carcass, 20-day adult    11 ovary, virgin 4-day female    21 ovary, mated 4-day female    24 testis, mated 4-day male    4 accessory gland, mated 4-day male    7 Expression Level Scale  Very low   Low   Moderate   Moderately high  Linear, scaled to Moderate expression Tissue   Expression Level imaginal disc, larvae L3 wandering    11 central nervous system, larvae L3    15 central nervous system, pupae P8  (37) head, virgin 1-day female    16 head, virgin 4-day female    16 head, virgin 20-day female    8 head, mated 1-day female    17 head, mated 4-day female    12 head, mated 20-day female    10 head, mated 1-day male    21 head, mated 4-day male    20 head, mated 20-day male    17 salivary gland, larvae L3 wandering    6 salivary gland, white prepupae    3 digestive system, larvae L3 wandering    6 digestive system, 1-day adult    3 digestive system, 4-day adult    5 digestive system, 20-day adult    3 fat body, larvae L3 wandering    9 fat body, white prepupae    13 fat body, pupae P8  (34) carcass, larvae L3 wandering    15 carcass, 1-day adult    14 carcass, 4-day adult    14 carcass, 20-day adult    11 ovary, virgin 4-day female    21 ovary, mated 4-day female    24 testis, mated 4-day male    4 accessory gland, mated 4-day male    7 Expression Level Scale  Very low   Low   Moderate   Moderately high  Linear, scaled to High expression Tissue   Expression Level imaginal disc, larvae L3 wandering    11 central nervous system, larvae L3    15 central nervous system, pupae P8    37 head, virgin 1-day female    16 head, virgin 4-day female    16 head, virgin 20-day female    8 head, mated 1-day female    17 head, mated 4-day female    12 head, mated 20-day female    10 head, mated 1-day male    21 head, mated 4-day male    20 head, mated 20-day male    17 salivary gland, larvae L3 wandering    6 salivary gland, white prepupae    3 digestive system, larvae L3 wandering    6 digestive system, 1-day adult    3 digestive system, 4-day adult    5 digestive system, 20-day adult    3 fat body, larvae L3 wandering    9 fat body, white prepupae    13 fat body, pupae P8    34 carcass, larvae L3 wandering    15 carcass, 1-day adult    14 carcass, 4-day adult    14 carcass, 20-day adult    11 ovary, virgin 4-day female    21 ovary, mated 4-day female    24 testis, mated 4-day male    4 accessory gland, mated 4-day male    7 Expression Level Scale  Very low   Low   Moderate   Moderately high   High   Very high  Linear, scaled to Extremely high expression Tissue   Expression Level imaginal disc, larvae L3 wandering    11 central nervous system, larvae L3    15 central nervous system, pupae P8    37 head, virgin 1-day female    16 head, virgin 4-day female    16 head, virgin 20-day female    8 head, mated 1-day female    17 head, mated 4-day female    12 head, mated 20-day female    10 head, mated 1-day male    21 head, mated 4-day male    20 head, mated 20-day male    17 salivary gland, larvae L3 wandering    6 salivary gland, white prepupae    3 digestive system, larvae L3 wandering    6 digestive system, 1-day adult    3 digestive system, 4-day adult    5 digestive system, 20-day adult    3 fat body, larvae L3 wandering    9 fat body, white prepupae    13 fat body, pupae P8    34 carcass, larvae L3 wandering    15 carcass, 1-day adult    14 carcass, 4-day adult    14 carcass, 20-day adult    11 ovary, virgin 4-day female    21 ovary, mated 4-day female    24 testis, mated 4-day male    4 accessory gland, mated 4-day male    7 Expression Level Scale  Extremely high  log, scaled to maximum expression level Tissue   Expression Level imaginal disc, larvae L3 wandering    11 central nervous system, larvae L3    15 central nervous system, pupae P8    37 head, virgin 1-day female    16 head, virgin 4-day female    16 head, virgin 20-day female    8 head, mated 1-day female    17 head, mated 4-day female    12 head, mated 20-day female    10 head, mated 1-day male    21 head, mated 4-day male    20 head, mated 20-day male    17 salivary gland, larvae L3 wandering    6 salivary gland, white prepupae    3 digestive system, larvae L3 wandering    6 digestive system, 1-day adult    3 digestive system, 4-day adult    5 digestive system, 20-day adult    3 fat body, larvae L3 wandering    9 fat body, white prepupae    13 fat body, pupae P8    34 carcass, larvae L3 wandering    15 carcass, 1-day adult    14 carcass, 4-day adult    14 carcass, 20-day adult    11 ovary, virgin 4-day female    21 ovary, mated 4-day female    24 testis, mated 4-day male    4 accessory gland, mated 4-day male    7 Expression Level Scale  Very low   Low   Moderate   Moderately high   High  log, scaled to Moderate expression Tissue   Expression Level imaginal disc, larvae L3 wandering    11 central nervous system, larvae L3    15 central nervous system, pupae P8  (37) head, virgin 1-day female    16 head, virgin 4-day female    16 head, virgin 20-day female    8 head, mated 1-day female    17 head, mated 4-day female    12 head, mated 20-day female    10 head, mated 1-day male    21 head, mated 4-day male    20 head, mated 20-day male    17 salivary gland, larvae L3 wandering    6 salivary gland, white prepupae    3 digestive system, larvae L3 wandering    6 digestive system, 1-day adult    3 digestive system, 4-day adult    5 digestive system, 20-day adult    3 fat body, larvae L3 wandering    9 fat body, white prepupae    13 fat body, pupae P8  (34) carcass, larvae L3 wandering    15 carcass, 1-day adult    14 carcass, 4-day adult    14 carcass, 20-day adult    11 ovary, virgin 4-day female    21 ovary, mated 4-day female    24 testis, mated 4-day male    4 accessory gland, mated 4-day male    7 Expression Level Scale  Very low   Low   Moderate   Moderately high  log, scaled to High expression Tissue   Expression Level imaginal disc, larvae L3 wandering    11 central nervous system, larvae L3    15 central nervous system, pupae P8    37 head, virgin 1-day female    16 head, virgin 4-day female    16 head, virgin 20-day female    8 head, mated 1-day female    17 head, mated 4-day female    12 head, mated 20-day female    10 head, mated 1-day male    21 head, mated 4-day male    20 head, mated 20-day male    17 salivary gland, larvae L3 wandering    6 salivary gland, white prepupae    3 digestive system, larvae L3 wandering    6 digestive system, 1-day adult    3 digestive system, 4-day adult    5 digestive system, 20-day adult    3 fat body, larvae L3 wandering    9 fat body, white prepupae    13 fat body, pupae P8    34 carcass, larvae L3 wandering    15 carcass, 1-day adult    14 carcass, 4-day adult    14 carcass, 20-day adult    11 ovary, virgin 4-day female    21 ovary, mated 4-day female    24 testis, mated 4-day male    4 accessory gland, mated 4-day male    7 Expression Level Scale  Very low   Low   Moderate   Moderately high   High   Very high  log, scaled to Extremely high expression Tissue   Expression Level imaginal disc, larvae L3 wandering    11 central nervous system, larvae L3    15 central nervous system, pupae P8    37 head, virgin 1-day female    16 head, virgin 4-day female    16 head, virgin 20-day female    8 head, mated 1-day female    17 head, mated 4-day female    12 head, mated 20-day female    10 head, mated 1-day male    21 head, mated 4-day male    20 head, mated 20-day male    17 salivary gland, larvae L3 wandering    6 salivary gland, white prepupae    3 digestive system, larvae L3 wandering    6 digestive system, 1-day adult    3 digestive system, 4-day adult    5 digestive system, 20-day adult    3 fat body, larvae L3 wandering    9 fat body, white prepupae    13 fat body, pupae P8    34 carcass, larvae L3 wandering    15 carcass, 1-day adult    14 carcass, 4-day adult    14 carcass, 20-day adult    11 ovary, virgin 4-day female    21 ovary, mated 4-day female    24 testis, mated 4-day male    4 accessory gland, mated 4-day male    7 Expression Level Scale  Very low   Low   Moderate   Moderately high   High   Very high   Extremely high  Heatmap Tissue   Expression Level imaginal disc, larvae L3 wandering     central nervous system, larvae L3     central nervous system, pupae P8     head, virgin 1-day female     head, virgin 4-day female     head, virgin 20-day female     head, mated 1-day female     head, mated 4-day female     head, mated 20-day female     head, mated 1-day male     head, mated 4-day male     head, mated 20-day male     salivary gland, larvae L3 wandering     salivary gland, white prepupae     digestive system, larvae L3 wandering     digestive system, 1-day adult     digestive system, 4-day adult     digestive system, 20-day adult     fat body, larvae L3 wandering     fat body, white prepupae     fat body, pupae P8     carcass, larvae L3 wandering     carcass, 1-day adult     carcass, 4-day adult     carcass, 20-day adult     ovary, virgin 4-day female     ovary, mated 4-day female     testis, mated 4-day male     accessory gland, mated 4-day male
modENCODE Development RNA-Seq
modENCODE Temporal Expression Data (modENCODE_mRNA-Seq_U)    Styles Linear Logarithmic Heatmap    Scales gene max expression Moderate expression bin max High expression bin max Extremely high expression bin max Summary of modENCODE Temporal Expression Profile: Temporal profile ranges from a peak of high expression to a trough of low expression. Peak expression observed during late pupal stages. [download data (TSV)] Guide to modENCODE expression level colors   No/Extremely low expression (0 - 0)   Very low expression (1 - 3)   Low expression (4 - 10)   Moderate expression (11 - 25)   Moderately high expression (26 - 50)   High expression (51 - 100)   Very high expression (101 - 1000)   Extremely high expression (>1000) Linear, scaled to maximum expression level Developmental Stage   Expression Level embryo 00-02hr    13 embryo 02-04hr    6 embryo 04-06hr    5 embryo 06-08hr    7 embryo 08-10hr    14 embryo 10-12hr    20 embryo 12-14hr    28 embryo 14-16hr    49 embryo 16-18hr    37 embryo 18-20hr    24 embryo 20-22hr    14 embryo 22-24hr    16 larva L1    9 larva L2    11 larva L3 12hr old    6 larva L3 puffstage 1-2    3 larva L3 puffstage 3-6    7 larva L3 puffstage 7-9    8 white prepupae new    8 white prepupae 12hr    18 white prepupae 24hr    24 pupae 2d postWPP    69 pupae 3d postWPP    46 pupae 4d postWPP    9 adult male 01day    10 adult male 05day    8 adult male 30day    6 adult female 01day    12 adult female 05day    14 adult female 30day    14 Expression Level Scale  Very low   Low   Moderate   Moderately high   High  Linear, scaled to Moderate expression Developmental Stage   Expression Level embryo 00-02hr    13 embryo 02-04hr    6 embryo 04-06hr    5 embryo 06-08hr    7 embryo 08-10hr    14 embryo 10-12hr    20 embryo 12-14hr  (28) embryo 14-16hr  (49) embryo 16-18hr  (37) embryo 18-20hr    24 embryo 20-22hr    14 embryo 22-24hr    16 larva L1    9 larva L2    11 larva L3 12hr old    6 larva L3 puffstage 1-2    3 larva L3 puffstage 3-6    7 larva L3 puffstage 7-9    8 white prepupae new    8 white prepupae 12hr    18 white prepupae 24hr    24 pupae 2d postWPP  (69) pupae 3d postWPP  (46) pupae 4d postWPP    9 adult male 01day    10 adult male 05day    8 adult male 30day    6 adult female 01day    12 adult female 05day    14 adult female 30day    14 Expression Level Scale  Very low   Low   Moderate   Moderately high  Linear, scaled to High expression Developmental Stage   Expression Level embryo 00-02hr    13 embryo 02-04hr    6 embryo 04-06hr    5 embryo 06-08hr    7 embryo 08-10hr    14 embryo 10-12hr    20 embryo 12-14hr    28 embryo 14-16hr    49 embryo 16-18hr    37 embryo 18-20hr    24 embryo 20-22hr    14 embryo 22-24hr    16 larva L1    9 larva L2    11 larva L3 12hr old    6 larva L3 puffstage 1-2    3 larva L3 puffstage 3-6    7 larva L3 puffstage 7-9    8 white prepupae new    8 white prepupae 12hr    18 white prepupae 24hr    24 pupae 2d postWPP    69 pupae 3d postWPP    46 pupae 4d postWPP    9 adult male 01day    10 adult male 05day    8 adult male 30day    6 adult female 01day    12 adult female 05day    14 adult female 30day    14 Expression Level Scale  Very low   Low   Moderate   Moderately high   High   Very high  Linear, scaled to Extremely high expression Developmental Stage   Expression Level embryo 00-02hr    13 embryo 02-04hr    6 embryo 04-06hr    5 embryo 06-08hr    7 embryo 08-10hr    14 embryo 10-12hr    20 embryo 12-14hr    28 embryo 14-16hr    49 embryo 16-18hr    37 embryo 18-20hr    24 embryo 20-22hr    14 embryo 22-24hr    16 larva L1    9 larva L2    11 larva L3 12hr old    6 larva L3 puffstage 1-2    3 larva L3 puffstage 3-6    7 larva L3 puffstage 7-9    8 white prepupae new    8 white prepupae 12hr    18 white prepupae 24hr    24 pupae 2d postWPP    69 pupae 3d postWPP    46 pupae 4d postWPP    9 adult male 01day    10 adult male 05day    8 adult male 30day    6 adult female 01day    12 adult female 05day    14 adult female 30day    14 Expression Level Scale  Extremely high  log, scaled to maximum expression level Developmental Stage   Expression Level embryo 00-02hr    13 embryo 02-04hr    6 embryo 04-06hr    5 embryo 06-08hr    7 embryo 08-10hr    14 embryo 10-12hr    20 embryo 12-14hr    28 embryo 14-16hr    49 embryo 16-18hr    37 embryo 18-20hr    24 embryo 20-22hr    14 embryo 22-24hr    16 larva L1    9 larva L2    11 larva L3 12hr old    6 larva L3 puffstage 1-2    3 larva L3 puffstage 3-6    7 larva L3 puffstage 7-9    8 white prepupae new    8 white prepupae 12hr    18 white prepupae 24hr    24 pupae 2d postWPP    69 pupae 3d postWPP    46 pupae 4d postWPP    9 adult male 01day    10 adult male 05day    8 adult male 30day    6 adult female 01day    12 adult female 05day    14 adult female 30day    14 Expression Level Scale  Very low   Low   Moderate   Moderately high   High   Very high  log, scaled to Moderate expression Developmental Stage   Expression Level embryo 00-02hr    13 embryo 02-04hr    6 embryo 04-06hr    5 embryo 06-08hr    7 embryo 08-10hr    14 embryo 10-12hr    20 embryo 12-14hr  28 embryo 14-16hr  (49) embryo 16-18hr  (37) embryo 18-20hr    24 embryo 20-22hr    14 embryo 22-24hr    16 larva L1    9 larva L2    11 larva L3 12hr old    6 larva L3 puffstage 1-2    3 larva L3 puffstage 3-6    7 larva L3 puffstage 7-9    8 white prepupae new    8 white prepupae 12hr    18 white prepupae 24hr    24 pupae 2d postWPP  (69) pupae 3d postWPP  (46) pupae 4d postWPP    9 adult male 01day    10 adult male 05day    8 adult male 30day    6 adult female 01day    12 adult female 05day    14 adult female 30day    14 Expression Level Scale  Very low   Low   Moderate   Moderately high  log, scaled to High expression Developmental Stage   Expression Level embryo 00-02hr    13 embryo 02-04hr    6 embryo 04-06hr    5 embryo 06-08hr    7 embryo 08-10hr    14 embryo 10-12hr    20 embryo 12-14hr    28 embryo 14-16hr    49 embryo 16-18hr    37 embryo 18-20hr    24 embryo 20-22hr    14 embryo 22-24hr    16 larva L1    9 larva L2    11 larva L3 12hr old    6 larva L3 puffstage 1-2    3 larva L3 puffstage 3-6    7 larva L3 puffstage 7-9    8 white prepupae new    8 white prepupae 12hr    18 white prepupae 24hr    24 pupae 2d postWPP    69 pupae 3d postWPP    46 pupae 4d postWPP    9 adult male 01day    10 adult male 05day    8 adult male 30day    6 adult female 01day    12 adult female 05day    14 adult female 30day    14 Expression Level Scale  Very low   Low   Moderate   Moderately high   High   Very high  log, scaled to Extremely high expression Developmental Stage   Expression Level embryo 00-02hr    13 embryo 02-04hr    6 embryo 04-06hr    5 embryo 06-08hr    7 embryo 08-10hr    14 embryo 10-12hr    20 embryo 12-14hr    28 embryo 14-16hr    49 embryo 16-18hr    37 embryo 18-20hr    24 embryo 20-22hr    14 embryo 22-24hr    16 larva L1    9 larva L2    11 larva L3 12hr old    6 larva L3 puffstage 1-2    3 larva L3 puffstage 3-6    7 larva L3 puffstage 7-9    8 white prepupae new    8 white prepupae 12hr    18 white prepupae 24hr    24 pupae 2d postWPP    69 pupae 3d postWPP    46 pupae 4d postWPP    9 adult male 01day    10 adult male 05day    8 adult male 30day    6 adult female 01day    12 adult female 05day    14 adult female 30day    14 Expression Level Scale  Very low   Low   Moderate   Moderately high   High   Very high   Extremely high  Heatmap Developmental Stage   Expression Level embryo 00-02hr     embryo 02-04hr     embryo 04-06hr     embryo 06-08hr     embryo 08-10hr     embryo 10-12hr     embryo 12-14hr     embryo 14-16hr     embryo 16-18hr     embryo 18-20hr     embryo 20-22hr     embryo 22-24hr     larva L1     larva L2     larva L3 12hr old     larva L3 puffstage 1-2     larva L3 puffstage 3-6     larva L3 puffstage 7-9     white prepupae new     white prepupae 12hr     white prepupae 24hr     pupae 2d postWPP     pupae 3d postWPP     pupae 4d postWPP     adult male 01day     adult male 05day     adult male 30day     adult female 01day     adult female 05day     adult female 30day     modENCODE Temporal Expression Data (Graveley et al., 2011)
modENCODE Cell Lines RNA-Seq
modENCODE Cell Line Expression Data (modENCODE_mRNA-Seq_cell.A) (modENCODE_mRNA-Seq_cell.B)    Styles Linear Logarithmic Heatmap    Scales gene max expression Moderate expression bin max High expression bin max Extremely high expression bin max [download data (TSV)] Guide to modENCODE expression level colors   No/Extremely low expression (0 - 0)   Very low expression (1 - 3)   Low expression (4 - 10)   Moderate expression (11 - 25)   Moderately high expression (26 - 50)   High expression (51 - 100)   Very high expression (101 - 1000)   Extremely high expression (>1000) Linear, scaled to maximum expression level Cell Line   Expression Level Schneider line 2 S2R+    1 Schneider line 2 Sg4    1 embryonic 1182-4H    2 embryonic GM2    2 embryonic Kc167    5 embryonic S1    2 embryonic S3    1 leg disc CME L1    1 wing disc CME-W2    0 wing disc ML-DmD8    1 wing disc ML-DmD9    0 wing disc ML-DmD16-c3    10 wing disc ML-DmD21    1 wing disc ML-DmD32    2 haltere disc ML-DmD17-c3    5 eye-antennal disc ML-DmD11    1 antennal disc ML-DmD20-c5    1 mixed discs ML-DmD4-c1    1 CNS ML-DmBG1-c1    1 CNS ML-DmBG2-c2    0 tumorous blood cells mbn2    1 ovary fGS/OSS    66 ovary OSC    44 ovary OSS    47 Expression Level Scale  Very low   Low   Moderate   Moderately high   High  Linear, scaled to Moderate expression Cell Line   Expression Level Schneider line 2 S2R+    1 Schneider line 2 Sg4    1 embryonic 1182-4H    2 embryonic GM2    2 embryonic Kc167    5 embryonic S1    2 embryonic S3    1 leg disc CME L1    1 wing disc CME-W2    0 wing disc ML-DmD8    1 wing disc ML-DmD9    0 wing disc ML-DmD16-c3    10 wing disc ML-DmD21    1 wing disc ML-DmD32    2 haltere disc ML-DmD17-c3    5 eye-antennal disc ML-DmD11    1 antennal disc ML-DmD20-c5    1 mixed discs ML-DmD4-c1    1 CNS ML-DmBG1-c1    1 CNS ML-DmBG2-c2    0 tumorous blood cells mbn2    1 ovary fGS/OSS  (66) ovary OSC  (44) ovary OSS  (47) Expression Level Scale  Very low   Low   Moderate   Moderately high  Linear, scaled to High expression Cell Line   Expression Level Schneider line 2 S2R+    1 Schneider line 2 Sg4    1 embryonic 1182-4H    2 embryonic GM2    2 embryonic Kc167    5 embryonic S1    2 embryonic S3    1 leg disc CME L1    1 wing disc CME-W2    0 wing disc ML-DmD8    1 wing disc ML-DmD9    0 wing disc ML-DmD16-c3    10 wing disc ML-DmD21    1 wing disc ML-DmD32    2 haltere disc ML-DmD17-c3    5 eye-antennal disc ML-DmD11    1 antennal disc ML-DmD20-c5    1 mixed discs ML-DmD4-c1    1 CNS ML-DmBG1-c1    1 CNS ML-DmBG2-c2    0 tumorous blood cells mbn2    1 ovary fGS/OSS    66 ovary OSC    44 ovary OSS    47 Expression Level Scale  Very low   Low   Moderate   Moderately high   High   Very high  Linear, scaled to Extremely high expression Cell Line   Expression Level Schneider line 2 S2R+    1 Schneider line 2 Sg4    1 embryonic 1182-4H    2 embryonic GM2    2 embryonic Kc167    5 embryonic S1    2 embryonic S3    1 leg disc CME L1    1 wing disc CME-W2    0 wing disc ML-DmD8    1 wing disc ML-DmD9    0 wing disc ML-DmD16-c3    10 wing disc ML-DmD21    1 wing disc ML-DmD32    2 haltere disc ML-DmD17-c3    5 eye-antennal disc ML-DmD11    1 antennal disc ML-DmD20-c5    1 mixed discs ML-DmD4-c1    1 CNS ML-DmBG1-c1    1 CNS ML-DmBG2-c2    0 tumorous blood cells mbn2    1 ovary fGS/OSS    66 ovary OSC    44 ovary OSS    47 Expression Level Scale  Extremely high  log, scaled to maximum expression level Cell Line   Expression Level Schneider line 2 S2R+    1 Schneider line 2 Sg4    1 embryonic 1182-4H    2 embryonic GM2    2 embryonic Kc167    5 embryonic S1    2 embryonic S3    1 leg disc CME L1    1 wing disc CME-W2    0 wing disc ML-DmD8    1 wing disc ML-DmD9    0 wing disc ML-DmD16-c3    10 wing disc ML-DmD21    1 wing disc ML-DmD32    2 haltere disc ML-DmD17-c3    5 eye-antennal disc ML-DmD11    1 antennal disc ML-DmD20-c5    1 mixed discs ML-DmD4-c1    1 CNS ML-DmBG1-c1    1 CNS ML-DmBG2-c2    0 tumorous blood cells mbn2    1 ovary fGS/OSS    66 ovary OSC    44 ovary OSS    47 Expression Level Scale  Very low   Low   Moderate   Moderately high   High  log, scaled to Moderate expression Cell Line   Expression Level Schneider line 2 S2R+    1 Schneider line 2 Sg4    1 embryonic 1182-4H    2 embryonic GM2    2 embryonic Kc167    5 embryonic S1    2 embryonic S3    1 leg disc CME L1    1 wing disc CME-W2    0 wing disc ML-DmD8    1 wing disc ML-DmD9    0 wing disc ML-DmD16-c3    10 wing disc ML-DmD21    1 wing disc ML-DmD32    2 haltere disc ML-DmD17-c3    5 eye-antennal disc ML-DmD11    1 antennal disc ML-DmD20-c5    1 mixed discs ML-DmD4-c1    1 CNS ML-DmBG1-c1    1 CNS ML-DmBG2-c2    0 tumorous blood cells mbn2    1 ovary fGS/OSS  (66) ovary OSC  (44) ovary OSS  (47) Expression Level Scale  Very low   Low   Moderate   Moderately high  log, scaled to High expression Cell Line   Expression Level Schneider line 2 S2R+    1 Schneider line 2 Sg4    1 embryonic 1182-4H    2 embryonic GM2    2 embryonic Kc167    5 embryonic S1    2 embryonic S3    1 leg disc CME L1    1 wing disc CME-W2    0 wing disc ML-DmD8    1 wing disc ML-DmD9    0 wing disc ML-DmD16-c3    10 wing disc ML-DmD21    1 wing disc ML-DmD32    2 haltere disc ML-DmD17-c3    5 eye-antennal disc ML-DmD11    1 antennal disc ML-DmD20-c5    1 mixed discs ML-DmD4-c1    1 CNS ML-DmBG1-c1    1 CNS ML-DmBG2-c2    0 tumorous blood cells mbn2    1 ovary fGS/OSS    66 ovary OSC    44 ovary OSS    47 Expression Level Scale  Very low   Low   Moderate   Moderately high   High   Very high  log, scaled to Extremely high expression Cell Line   Expression Level Schneider line 2 S2R+    1 Schneider line 2 Sg4    1 embryonic 1182-4H    2 embryonic GM2    2 embryonic Kc167    5 embryonic S1    2 embryonic S3    1 leg disc CME L1    1 wing disc CME-W2    0 wing disc ML-DmD8    1 wing disc ML-DmD9    0 wing disc ML-DmD16-c3    10 wing disc ML-DmD21    1 wing disc ML-DmD32    2 haltere disc ML-DmD17-c3    5 eye-antennal disc ML-DmD11    1 antennal disc ML-DmD20-c5    1 mixed discs ML-DmD4-c1    1 CNS ML-DmBG1-c1    1 CNS ML-DmBG2-c2    0 tumorous blood cells mbn2    1 ovary fGS/OSS    66 ovary OSC    44 ovary OSS    47 Expression Level Scale  Very low   Low   Moderate   Moderately high   High   Very high   Extremely high  Heatmap Cell Line   Expression Level Schneider line 2 S2R+     Schneider line 2 Sg4     embryonic 1182-4H     embryonic GM2     embryonic Kc167     embryonic S1     embryonic S3     leg disc CME L1     wing disc CME-W2     wing disc ML-DmD8     wing disc ML-DmD9     wing disc ML-DmD16-c3     wing disc ML-DmD21     wing disc ML-DmD32     haltere disc ML-DmD17-c3     eye-antennal disc ML-DmD11     antennal disc ML-DmD20-c5     mixed discs ML-DmD4-c1     CNS ML-DmBG1-c1     CNS ML-DmBG2-c2     tumorous blood cells mbn2     ovary fGS/OSS     ovary OSC     ovary OSS
modENCODE Treatments RNA-Seq
modENCODE Treatment Expression Data (modENCODE_mRNA-Seq_treatments)    Styles Linear Logarithmic Heatmap    Scales gene max expression Moderate expression bin max High expression bin max Extremely high expression bin max [download data (TSV)] Guide to modENCODE expression level colors   No/Extremely low expression (0 - 0)   Very low expression (1 - 3)   Low expression (4 - 10)   Moderate expression (11 - 25)   Moderately high expression (26 - 50)   High expression (51 - 100)   Very high expression (101 - 1000)   Extremely high expression (>1000) Linear, scaled to maximum expression level Treatment   Expression Level extended cold, 4-day adult    13 cold shock, 4-day adult    8 heat shock, 4-day adult    10 Cadmium 50 mM 6 hrs, larvae L3    2 Cadmium 50 mM 12 hrs, larvae L3    3 Cadmium 50 mM 48 hrs, 4-day adult    9 Cadmium 100 mM 48 hrs, 4-day adult    22 Copper 0.5 mM 12 hrs, larvae L3    5 Copper 15 mM 48 hrs, 4-day adult    27 Zinc 5 mM 12 hrs, larvae L3    3 Zinc 4.5 mM 48 hrs, 4-day adult    19 Ethanol 2.5% 3 hrs, larvae L3    17 Ethanol 5% 3 hrs, larvae L3    3 Ethanol 10% 3 hrs, larvae L3    2 Caffeine 1.5 mg/ml 4 hrs, larvae L3    3 Caffeine 2.5 mg/ml 48 hrs, 4-day adult    21 Caffeine 25 mg/ml 48 hrs, 4-day adult    13 Paraquat 5 mM 48 hrs, 4-day adult    16 Paraquat 10 mM 48 hrs, 4-day adult    15 Rotenone 2 μg 12 hrs, larvae L3    2 Rotenone 8 μg 12 hrs, larvae L3    2 Expression Level Scale  Very low   Low   Moderate   Moderately high  Linear, scaled to Moderate expression Treatment   Expression Level extended cold, 4-day adult    13 cold shock, 4-day adult    8 heat shock, 4-day adult    10 Cadmium 50 mM 6 hrs, larvae L3    2 Cadmium 50 mM 12 hrs, larvae L3    3 Cadmium 50 mM 48 hrs, 4-day adult    9 Cadmium 100 mM 48 hrs, 4-day adult    22 Copper 0.5 mM 12 hrs, larvae L3    5 Copper 15 mM 48 hrs, 4-day adult  (27) Zinc 5 mM 12 hrs, larvae L3    3 Zinc 4.5 mM 48 hrs, 4-day adult    19 Ethanol 2.5% 3 hrs, larvae L3    17 Ethanol 5% 3 hrs, larvae L3    3 Ethanol 10% 3 hrs, larvae L3    2 Caffeine 1.5 mg/ml 4 hrs, larvae L3    3 Caffeine 2.5 mg/ml 48 hrs, 4-day adult    21 Caffeine 25 mg/ml 48 hrs, 4-day adult    13 Paraquat 5 mM 48 hrs, 4-day adult    16 Paraquat 10 mM 48 hrs, 4-day adult    15 Rotenone 2 μg 12 hrs, larvae L3    2 Rotenone 8 μg 12 hrs, larvae L3    2 Expression Level Scale  Very low   Low   Moderate   Moderately high  Linear, scaled to High expression Treatment   Expression Level extended cold, 4-day adult    13 cold shock, 4-day adult    8 heat shock, 4-day adult    10 Cadmium 50 mM 6 hrs, larvae L3    2 Cadmium 50 mM 12 hrs, larvae L3    3 Cadmium 50 mM 48 hrs, 4-day adult    9 Cadmium 100 mM 48 hrs, 4-day adult    22 Copper 0.5 mM 12 hrs, larvae L3    5 Copper 15 mM 48 hrs, 4-day adult    27 Zinc 5 mM 12 hrs, larvae L3    3 Zinc 4.5 mM 48 hrs, 4-day adult    19 Ethanol 2.5% 3 hrs, larvae L3    17 Ethanol 5% 3 hrs, larvae L3    3 Ethanol 10% 3 hrs, larvae L3    2 Caffeine 1.5 mg/ml 4 hrs, larvae L3    3 Caffeine 2.5 mg/ml 48 hrs, 4-day adult    21 Caffeine 25 mg/ml 48 hrs, 4-day adult    13 Paraquat 5 mM 48 hrs, 4-day adult    16 Paraquat 10 mM 48 hrs, 4-day adult    15 Rotenone 2 μg 12 hrs, larvae L3    2 Rotenone 8 μg 12 hrs, larvae L3    2 Expression Level Scale  Very low   Low   Moderate   Moderately high   High   Very high  Linear, scaled to Extremely high expression Treatment   Expression Level extended cold, 4-day adult    13 cold shock, 4-day adult    8 heat shock, 4-day adult    10 Cadmium 50 mM 6 hrs, larvae L3    2 Cadmium 50 mM 12 hrs, larvae L3    3 Cadmium 50 mM 48 hrs, 4-day adult    9 Cadmium 100 mM 48 hrs, 4-day adult    22 Copper 0.5 mM 12 hrs, larvae L3    5 Copper 15 mM 48 hrs, 4-day adult    27 Zinc 5 mM 12 hrs, larvae L3    3 Zinc 4.5 mM 48 hrs, 4-day adult    19 Ethanol 2.5% 3 hrs, larvae L3    17 Ethanol 5% 3 hrs, larvae L3    3 Ethanol 10% 3 hrs, larvae L3    2 Caffeine 1.5 mg/ml 4 hrs, larvae L3    3 Caffeine 2.5 mg/ml 48 hrs, 4-day adult    21 Caffeine 25 mg/ml 48 hrs, 4-day adult    13 Paraquat 5 mM 48 hrs, 4-day adult    16 Paraquat 10 mM 48 hrs, 4-day adult    15 Rotenone 2 μg 12 hrs, larvae L3    2 Rotenone 8 μg 12 hrs, larvae L3    2 Expression Level Scale  Extremely high  log, scaled to maximum expression level Treatment   Expression Level extended cold, 4-day adult    13 cold shock, 4-day adult    8 heat shock, 4-day adult    10 Cadmium 50 mM 6 hrs, larvae L3    2 Cadmium 50 mM 12 hrs, larvae L3    3 Cadmium 50 mM 48 hrs, 4-day adult    9 Cadmium 100 mM 48 hrs, 4-day adult    22 Copper 0.5 mM 12 hrs, larvae L3    5 Copper 15 mM 48 hrs, 4-day adult    27 Zinc 5 mM 12 hrs, larvae L3    3 Zinc 4.5 mM 48 hrs, 4-day adult    19 Ethanol 2.5% 3 hrs, larvae L3    17 Ethanol 5% 3 hrs, larvae L3    3 Ethanol 10% 3 hrs, larvae L3    2 Caffeine 1.5 mg/ml 4 hrs, larvae L3    3 Caffeine 2.5 mg/ml 48 hrs, 4-day adult    21 Caffeine 25 mg/ml 48 hrs, 4-day adult    13 Paraquat 5 mM 48 hrs, 4-day adult    16 Paraquat 10 mM 48 hrs, 4-day adult    15 Rotenone 2 μg 12 hrs, larvae L3    2 Rotenone 8 μg 12 hrs, larvae L3    2 Expression Level Scale  Very low   Low   Moderate   Moderately high  log, scaled to Moderate expression Treatment   Expression Level extended cold, 4-day adult    13 cold shock, 4-day adult    8 heat shock, 4-day adult    10 Cadmium 50 mM 6 hrs, larvae L3    2 Cadmium 50 mM 12 hrs, larvae L3    3 Cadmium 50 mM 48 hrs, 4-day adult    9 Cadmium 100 mM 48 hrs, 4-day adult    22 Copper 0.5 mM 12 hrs, larvae L3    5 Copper 15 mM 48 hrs, 4-day adult  27 Zinc 5 mM 12 hrs, larvae L3    3 Zinc 4.5 mM 48 hrs, 4-day adult    19 Ethanol 2.5% 3 hrs, larvae L3    17 Ethanol 5% 3 hrs, larvae L3    3 Ethanol 10% 3 hrs, larvae L3    2 Caffeine 1.5 mg/ml 4 hrs, larvae L3    3 Caffeine 2.5 mg/ml 48 hrs, 4-day adult    21 Caffeine 25 mg/ml 48 hrs, 4-day adult    13 Paraquat 5 mM 48 hrs, 4-day adult    16 Paraquat 10 mM 48 hrs, 4-day adult    15 Rotenone 2 μg 12 hrs, larvae L3    2 Rotenone 8 μg 12 hrs, larvae L3    2 Expression Level Scale  Very low   Low   Moderate   Moderately high  log, scaled to High expression Treatment   Expression Level extended cold, 4-day adult    13 cold shock, 4-day adult    8 heat shock, 4-day adult    10 Cadmium 50 mM 6 hrs, larvae L3    2 Cadmium 50 mM 12 hrs, larvae L3    3 Cadmium 50 mM 48 hrs, 4-day adult    9 Cadmium 100 mM 48 hrs, 4-day adult    22 Copper 0.5 mM 12 hrs, larvae L3    5 Copper 15 mM 48 hrs, 4-day adult    27 Zinc 5 mM 12 hrs, larvae L3    3 Zinc 4.5 mM 48 hrs, 4-day adult    19 Ethanol 2.5% 3 hrs, larvae L3    17 Ethanol 5% 3 hrs, larvae L3    3 Ethanol 10% 3 hrs, larvae L3    2 Caffeine 1.5 mg/ml 4 hrs, larvae L3    3 Caffeine 2.5 mg/ml 48 hrs, 4-day adult    21 Caffeine 25 mg/ml 48 hrs, 4-day adult    13 Paraquat 5 mM 48 hrs, 4-day adult    16 Paraquat 10 mM 48 hrs, 4-day adult    15 Rotenone 2 μg 12 hrs, larvae L3    2 Rotenone 8 μg 12 hrs, larvae L3    2 Expression Level Scale  Very low   Low   Moderate   Moderately high   High   Very high  log, scaled to Extremely high expression Treatment   Expression Level extended cold, 4-day adult    13 cold shock, 4-day adult    8 heat shock, 4-day adult    10 Cadmium 50 mM 6 hrs, larvae L3    2 Cadmium 50 mM 12 hrs, larvae L3    3 Cadmium 50 mM 48 hrs, 4-day adult    9 Cadmium 100 mM 48 hrs, 4-day adult    22 Copper 0.5 mM 12 hrs, larvae L3    5 Copper 15 mM 48 hrs, 4-day adult    27 Zinc 5 mM 12 hrs, larvae L3    3 Zinc 4.5 mM 48 hrs, 4-day adult    19 Ethanol 2.5% 3 hrs, larvae L3    17 Ethanol 5% 3 hrs, larvae L3    3 Ethanol 10% 3 hrs, larvae L3    2 Caffeine 1.5 mg/ml 4 hrs, larvae L3    3 Caffeine 2.5 mg/ml 48 hrs, 4-day adult    21 Caffeine 25 mg/ml 48 hrs, 4-day adult    13 Paraquat 5 mM 48 hrs, 4-day adult    16 Paraquat 10 mM 48 hrs, 4-day adult    15 Rotenone 2 μg 12 hrs, larvae L3    2 Rotenone 8 μg 12 hrs, larvae L3    2 Expression Level Scale  Very low   Low   Moderate   Moderately high   High   Very high   Extremely high  Heatmap Treatment   Expression Level extended cold, 4-day adult     cold shock, 4-day adult     heat shock, 4-day adult     Cadmium 50 mM 6 hrs, larvae L3     Cadmium 50 mM 12 hrs, larvae L3     Cadmium 50 mM 48 hrs, 4-day adult     Cadmium 100 mM 48 hrs, 4-day adult     Copper 0.5 mM 12 hrs, larvae L3     Copper 15 mM 48 hrs, 4-day adult     Zinc 5 mM 12 hrs, larvae L3     Zinc 4.5 mM 48 hrs, 4-day adult     Ethanol 2.5% 3 hrs, larvae L3     Ethanol 5% 3 hrs, larvae L3     Ethanol 10% 3 hrs, larvae L3     Caffeine 1.5 mg/ml 4 hrs, larvae L3     Caffeine 2.5 mg/ml 48 hrs, 4-day adult     Caffeine 25 mg/ml 48 hrs, 4-day adult     Paraquat 5 mM 48 hrs, 4-day adult     Paraquat 10 mM 48 hrs, 4-day adult     Rotenone 2 μg 12 hrs, larvae L3     Rotenone 8 μg 12 hrs, larvae L3
Expression Clusters
	A cluster of genes with similar mRNA expression dynamics across development. (The modENCODE Consortium, 2010) mE2_34_mRNA_expression_cluster_09
External Data & Images
Linkouts	BDGP expression data • LD14119 FLIGHT - Cell culture data for RNAi and other high-throughput technologies • FBgn0003016 FlyAtlas - Adult expression by tissue, using Affymetrix Dros2 array • CG3479-RA FlyExpress - Embryonic expression images (BDGP data) • Show all FlyExpress processed images for osp Stages(s) 11-12 Stages(s) 13-16
Alleles & Phenotypes
Summary of Allele Phenotypes
Lethality Allele lethal, with Scer\GAL4Act5C.PU ospGD1504 lethal | recessive osprJ571 partially lethal - majority die osprJ571 partially lethal - majority die | recessive osprJ571 viable osp1 osppb3 Sterility Allele fertile osp1 Other Phenotypes Allele eye color defective, with Scer\GAL4GMR.PF, Scer\GAL4ey.PH ospGD1504 visible osp1 osp76e ospA5 ospalw visible | recessive osprJ571 Phenotype manifest in Allele crystal cell osp29 osprJ571 thorax, with Scer\GAL4pnr-MD237 ospGD1504 wing osp1 osp76e ospalw osprJ571 ospA5 wing, with Scer\GAL4Bx-MS1096 ospGD1504 wing (with Df(2L)A72) ospA5 wing (with Df(2L)A263) ospA5 wing (with Df(2L)osp29) ospA5 wing (with osp1) ospA5 wing (with ospA5) osp1 osprJ571 wing (with osprJ571) ospA5
Classical Alleles ( 56 )
For All Classical Alleles Show
Allele of osp Class Mutagen Stocks Known lesion osp29 gamma ray 2 -- ospKG06771 2 -- osp00865 P-element activity 1 -- osp1 ethyl methanesulfonate 1 -- osp22 ethyl methanesulfonate 1 -- osp76e ethyl methanesulfonate 1 -- ospB307 1 -- ospc00145 piggyBac transposase 1 -- ospc04765 piggyBac transposase 1 -- ospc05698 piggyBac transposase 1 -- ospCB-5301-3 1 -- ospd06776 P-element activity 1 -- ospd11647 P-element activity 1 -- ospe02984 piggyBac transposase 1 -- ospe02985 piggyBac transposase 1 -- ospf01299 1 -- ospf03217 piggyBac transposase 1 -- ospf05495 piggyBac transposase 1 -- ospf07101 piggyBac transposase 1 -- ospk13218 P-element activity 1 Yes ospMI00193 1 -- ospMI03036 1 -- ospMI05112 1 -- ospNP6350 1 -- osprJ571 P-element activity 1 Yes osp104 gamma ray 0 -- osp16 ethyl methanesulfonate 0 -- osp19 ethyl methanesulfonate 0 -- osp201 gamma ray 0 -- osp24 ethyl methanesulfonate 0 -- osp25 ethyl methanesulfonate 0 -- osp27 ethyl methanesulfonate 0 -- osp35 gamma ray 0 -- osp3 gamma ray 0 -- osp40 gamma ray 0 -- osp4 ethyl methanesulfonate 0 -- osp59 gamma ray 0 -- osp6 ethyl methanesulfonate 0 -- osp77e ethyl methanesulfonate 0 -- osp7 ethyl methanesulfonate 0 -- osp80.1D diepoxyoctane 0 -- osp83 gamma ray 0 -- osp8 ethyl methanesulfonate 0 -- osp90 gamma ray 0 -- osp91g2 ethyl methanesulfonate 0 -- ospA5 0 Yes ospalw 0 -- ospBG01265 P-element activity 0 -- ospCH62 ethyl methanesulfonate 0 -- ospCH63 ethyl methanesulfonate 0 -- ospf05599 piggyBac transposase 0 -- ospH47 alpha ray 0 -- ospH86 0 -- ospMpe X ray 0 -- osppb3 X ray 0 -- ospPW P-element activity 0 --
Alleles Carried on Transgenic Constructs ( 3 )
For All Alleles Carried on Transgenic Constructs Show
Allele of osp Class Mutagen Stocks Known lesion ospGD1504 in vitro construct - RNAi in vitro construct - regulatory fusion 1 Yes ospKK101464 in vitro construct - RNAi in vitro construct - regulatory fusion 0 Yes ospNIG.3479R in vitro construct - RNAi in vitro construct - regulatory fusion 0 Yes
Aneuploid Aberrations
Disrupted in	Df(2L)64j (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979) Df(2L)A47 (Ashburner et al., 1982, Gubb et al., 1990, Fossett et al., 1994, McNabb et al., 1996, Roote et al., 1996) Df(2L)A48 (Ashraf et al., 1999, Ashburner et al., 1982, Ashburner et al., 1982, Gubb et al., 1990, Fossett et al., 1994, McNabb et al., 1996, Roote et al., 1996, Grau et al., 1984) Df(2L)A63 (Ashburner et al., 1982, Ashburner et al., 1982, Fossett et al., 1994, McNabb et al., 1996, Roote et al., 1996, Roote, 1996.8) Df(2L)A72 (Ashburner et al., 1982, Ashburner et al., 1982, Fossett et al., 1994, McNabb et al., 1996, Roote et al., 1996) Df(2L)A178 (Ashburner et al., 1982, Ashburner et al., 1982, Fossett et al., 1994, Roote et al., 1996) Df(2L)A215 (Ashburner et al., 1982, Fossett et al., 1994) Df(2L)A217 (Ashburner et al., 1982, Fossett et al., 1994, McNabb et al., 1996, Roote, 1996.8) Df(2L)A220 (Ashburner et al., 1982, Fossett et al., 1994, McNabb et al., 1996, Roote et al., 1996) Df(2L)A245 (Ashburner et al., 1982, Fossett et al., 1994, McNabb et al., 1996, Roote et al., 1996, Roote, 1996.8) Df(2L)A246 (Ashburner et al., 1982, Fossett et al., 1994) Df(2L)A260 (Ashburner et al., 1982, McNabb et al., 1996, Roote, 1996.8) Df(2L)A263 (Ashburner et al., 1982) Df(2L)A264 (Ashburner et al., 1982, Fossett et al., 1994) Df(2L)A266 (Ashburner et al., 1982, Ashburner et al., 1982, Fossett et al., 1994, McNabb et al., 1996, Roote, 1996.8) Df(2L)A267 (Ashburner et al., 1982, Ashburner et al., 1982, Fossett et al., 1994, Roote et al., 1996) Df(2L)A337 (Fossett et al., 1994) Df(2L)A376 (Ashburner et al., 1982, Fossett et al., 1994) Df(2L)A377 (Ashburner et al., 1982) Df(2L)A379 (Ashburner et al., 1982, Ashburner et al., 1982, Fossett et al., 1994, McNabb et al., 1996) Df(2L)A400 (Ashburner et al., 1982, Ashburner et al., 1982, Fossett et al., 1994, Roote et al., 1996) Df(2L)A445 (Ashburner et al., 1982, Fossett et al., 1994, McNabb et al., 1996, Roote et al., 1996, Roote, 1996.8) Df(2L)Adh-nBR41 (Fossett et al., 1990) Df(2L)Adh-nBR55 (Fossett et al., 1990) Df(2L)Adh-nBR100 (Fossett et al., 1994) Df(2L)Adh-nBR101 (Fossett et al., 1994) Df(2L)Adh-nBR102 (Fossett et al., 1994, Roote et al., 1996) Df(2L)Adh-nBR103 (Fossett et al., 1994) Df(2L)Adh-nBR105 (Fossett et al., 1994, Roote, 1996.1.11, Roote et al., 1996, Roote, 1996.8) Df(2L)Adh-nBR106 (Fossett et al., 1994) Df(2L)Adh-nBR107 (Fossett et al., 1994, Roote, 1996.1.11, Roote et al., 1996, Roote, 1996.8) Df(2L)Adh-nBR111 (Fossett et al., 1990) Df(2L)Adh-nBR117 (Fossett et al., 1994) Df(2L)Adh-nBR118 (Fossett et al., 1994) Df(2L)Adh-nBR119 (Fossett et al., 1994) Df(2L)Adh-nBR120 (Fossett et al., 1994) Df(2L)Adh-nBR121 (Fossett et al., 1994, Roote, 1996.1.11, Roote et al., 1996, Roote, 1996.8) Df(2L)Adh-nBR122 (Fossett et al., 1994, Roote, 1996.1.11, Roote et al., 1996, Roote, 1996.8) Df(2L)Adh-nBR125 (Fossett et al., 1994) Df(2L)Adh-nBR126 (Fossett et al., 1994) Df(2L)Adh-nBR127 (Fossett et al., 1994, Roote, 1996.1.11, Roote et al., 1996, Roote, 1996.8) Df(2L)Adh-nBR128 (Fossett et al., 1994, Roote, 1996.1.11, Roote et al., 1996, Roote, 1996.8) Df(2L)Adh-nBR129 (Fossett et al., 1994) Df(2L)Adh-nBR130 (Fossett et al., 1994) Df(2L)Adh-nBR131 (Fossett et al., 1994) Df(2L)ARR1 (Roote et al., 1996) Df(2L)b84a1 (McNabb et al., 1996, Roote, 1996.8) Df(2L)b84h50 (Spradling et al., 1999, BDGP Project Members, 1994-1999) Df(2L)b88c25 (Roote, 2000.6.5, Roote, 2000.7.3, Roote, 2002.1.15) Df(2L)b89e68 (Roote, 2000.6.5, Roote, 2000.7.3, Roote, 2002.1.15) Df(2L)b89e88b (Roote, 2000.6.5) Df(2L)do1 (Ashburner et al., 1981) Df(2L)ED3 (Roote, 2002.2.6, Ryder and Roote, 2004.11.10) Df(2L)el77 (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979, McNabb et al., 1996) Df(2L)el80f1 (Ashburner et al., 1983) Df(2L)el81i1 (Roote, 2002.1.15) Df(2L)Exel8033 (Ryder, 2004.4.26) Df(2L)fn1 (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979, Ashburner et al., 1982, Ashburner et al., 1982, Grau et al., 1984) Df(2L)fn2 (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979, Ashburner et al., 1982, Ashburner et al., 1982, Grau et al., 1984, McNabb et al., 1996, Roote et al., 1996, Roote, 1996.8) Df(2L)fn3 (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979, Ashburner et al., 1982) Df(2L)fn5 (Ashburner et al., 1982) Df(2L)fn7 (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979, Ashburner et al., 1982, McNabb et al., 1996, Roote, 1996.8) Df(2L)fn12 (Woodruff and Ashburner, 1979, Ashburner et al., 1982) Df(2L)fn15 (Ashburner et al., 1982) Df(2L)fn26 (Ashburner et al., 1982) Df(2L)fn27 (Ashburner et al., 1982, Ashburner et al., 1982) Df(2L)fn30 (Ashburner et al., 1982) Df(2L)fn31 (Ashburner et al., 1982) Df(2L)fn36 (Ashburner et al., 1982, McNabb et al., 1996, Roote, 1996.8) Df(2L)fn52 (McNabb et al., 1996, Roote et al., 1996, Roote, 1996.8) Df(2L)GT5 (Gubb et al., 1990) Df(2L)k08808-mr1 (Roote, 2002.2.18) Df(2L)n78l3 (Ashburner et al., 1982) Df(2L)nNxF1 (Chia et al., 1985) Df(2L)nNxF2 (Chia et al., 1985) Df(2L)noc10 (McNabb et al., 1996, Roote, 1996.8) Df(2L)noc13 (McNabb et al., 1996, Roote, 1996.8) Df(2L)noc20 (McNabb et al., 1996) Df(2L)osp18 (Ashraf et al., 1999, McNabb et al., 1996, Roote et al., 1996) Df(2L)osp29 (Ashraf et al., 1999, McNabb et al., 1996, Roote et al., 1996, Roote, 1996.8) Df(2L)osp144 (Ashburner et al., 1982, McNabb et al., 1996, Roote et al., 1996, Roote, 1996.8) Df(2L)osp201 (McNabb et al., 1996) Df(2L)osp-204   Df(2L)PA4 (Roote, 2002.1.15, Ashburner et al., 1983) Df(2L)Sco-2   Df(2L)Sco7 (Ashburner et al., 1983) Df(2L)Scorv4 (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979, Ashburner et al., 1983, Ashburner et al., 1990) Df(2L)Scorv10 (Woodruff and Ashburner, 1979, Ashburner et al., 1980, Ashburner et al., 1983, Grau et al., 1984) Df(2L)Scorv14 (Woodruff and Ashburner, 1979, Ashburner et al., 1980, Ashburner et al., 1983) Df(2L)Scorv18 (Ashburner et al., 1980, Ashburner et al., 1983) Df(2L)Scorv23 (Ashburner et al., 1983, Grau et al., 1984) Df(2L)TE35B-1 (Gubb et al., 1985) Df(2L)TE35B-2 (Gubb et al., 1985) Df(2L)TE35B-3 (Gubb et al., 1985) Df(2L)TE35B-5 (Gubb et al., 1985) Df(2L)TE35B-6 (Gubb et al., 1985) Df(2L)TE35B-7 (Gubb et al., 1985, Roote et al., 1996) Df(2L)TE35B-8 (Gubb et al., 1985, McNabb et al., 1996, Roote et al., 1996) Df(2L)TE35B-9 (Gubb et al., 1985, McNabb et al., 1996, Roote, 1996.8) Df(2L)TE35B-11 (Gubb et al., 1985, McNabb et al., 1996, Roote et al., 1996, Roote, 1996.8) Df(2L)TE35B-15 (Gubb et al., 1985, McNabb et al., 1996, Roote, 1996.8) Df(2L)TE35B-48 (Gubb et al., 1986, McNabb et al., 1996, Roote, 1996.8, Gubb et al., 1997) Df(2L)TE35B-54 (Gubb et al., 1986, McNabb et al., 1996, Gubb et al., 1997) Df(2L)TE35B-54L (Roote et al., 1996) Df(2L)TE35BC-6 (Gubb et al., 1984) Df(2L)TE35BC-7 (Gubb et al., 1984) Df(2L)TE35BC-8 (Gubb et al., 1984) Df(2L)TE35BC-23 (Gubb et al., 1984) Df(2L)TE35BC-29 (Gubb et al., 1984) Df(2L)TE35BC-31 (Gubb et al., 1984) Df(2L)W (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979, Ashburner et al., 1982, McNabb et al., 1996, Roote, 1996.8) In(2L)75c (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979, Grau et al., 1984) In(2L)75cLC158.1R (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979) In(2L)b88e16 (Roote, 2000.6.5) In(2L)C158LScorv11R (Ashburner et al., 1983) In(2L)C158LScorv17R (Ashburner et al., 1983) In(2L)dppH86 (Roote et al., 1996) In(2L)osp22 (McNabb et al., 1996, Roote et al., 1996) In(2L)osp59 (McNabb et al., 1996, Roote et al., 1996) In(2L)TE35B-13 (Gubb et al., 1985) In(2LR)el6LA379R (Roote, 2002.1.15, Roote, 1996.1.11, Roote et al., 1996, Roote, 1996.8) In(2LR)noc4LScorv9R (Ashburner and Harrington, 1984) In(2LR)Scorv1LTE35B-15R (Roote et al., 1996) In(2LR)TE35B-4LScorv1R (Roote, 2002.1.15) In(2LR)TE35B-6LScorv9R (Ashburner and Harrington, 1984) In(2LR)TE35B-14LScorv1R (Roote, 2002.1.15) T(2;3)88g68 (Roote, 2000.6.5) T(2;3)GT7 (Gubb et al., 1990) T(2;3)H47 (McNabb et al., 1996, Roote et al., 1996) T(2;3)Mpe (McNabb et al., 1996) T(2;3)osp90 (McNabb et al., 1996, Roote et al., 1996) T(2;3)Scorv7 (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979, Ashburner et al., 1980, Ashburner et al., 1983, Ashburner and Harrington, 1984, Grau et al., 1984) T(2;3)TE35B-2a (Chia et al., 1985) T(2;3)TE35B-3DG16P (Chia et al., 1985) T(Y;2)A80DR15P (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979, Ashburner et al., 1982) T(Y;2)GT1 (Gubb et al., 1990) Tp(1;2)TE35B-ospA5 (Moschetti et al., 2004) Tp(2;2)A446 (Ashburner et al., 1982, Ashburner et al., 1982, Fossett et al., 1994, Roote et al., 1996) Tp(2;3)osp3 (McNabb et al., 1996) Tp(2;3)pb3 (Ashburner et al., 1981, McNabb et al., 1996, Roote et al., 1996) Tp(3;2)osp204 (Roote et al., 1996) Ts(2Lt;3Lt)GT7+Ts(2Rt;3Rt)Scorv13 (Roote, 2002.1.15, Roote et al., 1996) Ts(2Lt;3Lt)Mpe+Ts(2Rt;3Rt)G16 (Chia et al., 1985) Ts(2Lt;3Lt)TE35B-4+Ts(2Rt;3Rt)Scorv13 (Ashburner and Harrington, 1984, Chia et al., 1985) Ts(2Lt;3Lt)TE35B-28+Ts(2Rt;3Rt)osp90 (Roote, 1995.3.27, Roote et al., 1996) Ts(2Lt;3Lt)TE35BC-3+Ts(2Rt;3Rt)Scorv13 (Ashburner et al., 1983) Ts(2Lt;4Lt)TE35B-50+Ts(2Rt;4Rt)DTD22 (Roote, 2002.1.15) Ts(2Lt;4Lt)TE35B-101+Ts(2Rt;4Rt)DTD22 (Roote, 2002.1.15) Ts(Y;2Lt)el4+Ts(YSt;2Rt)R15 (Ashburner et al., 1982, Roote et al., 1996) Ts(Y;2Lt)GT1+Ts(YSt;2Rt)R15 (Roote et al., 1996) Ts(YLt;2Lt)G74+Ts(YSt;2Rt)B210 (Woodruff and Ashburner, 1979) Ts(YSt;2Lt)G74+Ts(YSt;2Rt)J165 (Woodruff and Ashburner, 1979) Ts(YSt;2Lt)G74+Ts(YSt;2Rt)P58 (Woodruff and Ashburner, 1979) Ts(YSt;2Lt)G74+Ts(YSt;2Rt)R15 (Woodruff and Ashburner, 1979)
Not disrupted in	Df(2L)A48 (Spradling et al., 1999, BDGP Project Members, 1994-1999) Df(2L)b75 (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979, Gubb et al., 1990) Df(2L)b80e3 (Ashburner et al., 1982, Roote et al., 1996) Df(2L)b81a1 (Ashburner et al., 1982, Roote et al., 1996) Df(2L)b83d29a (Roote et al., 1996) Df(2L)b84a2 (Roote et al., 1996) Df(2L)b84a4 (Roote, 2000.7.3, McNabb et al., 1996, Roote et al., 1996) Df(2L)b84a8 (Roote, 2000.7.3, McNabb et al., 1996) Df(2L)b88h49 (Roote et al., 1996) Df(2L)el14 (Roote et al., 1996) Df(2L)GT4 (Gubb et al., 1990, McNabb et al., 1996) Df(2L)osp29 (Ashburner et al., 1982, Chia et al., 1985, Grau et al., 1984) Df(2L)Sco-1 (Ashburner, 1982) Df(2L)Scorv16 (Ashburner et al., 1983, Grau et al., 1984) Df(2L)Scorv19 (Ashburner et al., 1983, Grau et al., 1984) Df(2L)Scorv25 (Ashburner et al., 1983, Grau et al., 1984) Df(2L)SS1 (Grau et al., 1984) Df(2L)TE35B-4 (Gubb et al., 1985) Df(2L)TE35B-10 (Gubb et al., 1985) Df(2L)TE35B-12 (Gubb et al., 1985, Roote et al., 1996, Roote, 1996.8) Df(2L)TE35B-203 (Roote, 2002.1.15) Df(2L)TE35B-300 (Roote et al., 1996) Df(2L)TE35BC-1 (Gubb et al., 1984) Df(2L)TE35BC-3 (Ashburner et al., 1982, Gubb et al., 1984, Grau et al., 1984) Df(2L)TE35BC-4 (Ashburner et al., 1982, Gubb et al., 1984, McNabb et al., 1996, Roote et al., 1996) Df(2L)TE35BC-24 (Gubb et al., 1984) Df(2L)TE35BC-28 (Gubb et al., 1984, McNabb et al., 1996) Df(2L)TE35BC-34 (Gubb et al., 1984, Roote, 1996.8) Df(2L)TE35BC-35 (Gubb et al., 1984, McNabb et al., 1996) Dp(2;2)Cam12 (Roote et al., 1996) In(2L)C158 (McNabb et al., 1996, Roote et al., 1996) In(2L)C158LScorv11R (Ashburner et al., 1981, Ashburner and Harrington, 1984) In(2L)C158LScorv17R (Ashburner and Harrington, 1984) In(2L)Scorv11 (Roote et al., 1996) In(2L)Scorv17 (Roote et al., 1996) In(2LR)b81a2LDTD43R (Roote, 2000.7.3, Roote, 2002.1.15) In(2LR)D20Lnoc4R (Roote et al., 1996) In(2LR)noc4LTE35B-50R (Roote, 2002.1.15) T(2;3)GT7 (Roote et al., 1996) T(2;3)TE35BC-3 (McNabb et al., 1996, Roote et al., 1996) T(Y;2)GT1 (Roote et al., 1996) T(Y;2)J165DP58P (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979) T(Y;2)R15DJ165P (Woodruff and Ashburner, 1979, Woodruff and Ashburner, 1979) Ts(2Lt;3Lt)dpps19+Ts(2Rt;3Rt)el24 (Roote et al., 1996) Ts(2Lt;3Lt)H118+Ts(2Rt;3Rt)TE35B-8 (Roote, 2002.1.15) Ts(2Lt;3Lt)H147+Ts(2Rt;3Rt)TE35B-8 (Roote, 2002.1.15) Ts(2Lt;3Lt)TE35B-4+Ts(2Rt;3Rt)Scorv13 (Roote, 2002.1.15) Ts(2Lt;3Lt)TE35BC-3+Ts(2Rt;3Rt)Scorv13 (Ashburner and Harrington, 1984, McNabb et al., 1996) Ts(2Lt;3Rt)GT10+Ts(2Rt;3Lt)TE35B-209 (Roote et al., 1996) Ts(2Lt;4Lt)GT6+Ts(2Rt;4Rt)TE35B-101 (Roote, 2002.1.15) Ts(Y;2Lt)el4+Ts(YSt;2Rt)A80 (Roote et al., 1996) Ts(Y;2Lt)GT2+Ts(YSt;2Rt)A80 (Roote et al., 1996) Ts(YSt;2Lt)G74+Ts(YSt;2Rt)A80 (Woodruff and Ashburner, 1979)
Not duplicated in	In(2L)C163.41LC158R (Ashburner et al., 1982, Roote et al., 1996) In(2LR)TE35B-226LDTD128R (Roote et al., 1996) Ts(2Lt;4Lt)TE35B-101+Ts(2Rt;4Rt)GT6 (Roote et al., 1996) Ts(Y;2Lt)TE35B-18+Ts(YSt;2Rt)A80 (Roote et al., 1996) Ts(YLt;2Lt)A80+Ts(Y;2Rt)el4 (Roote et al., 1996) Ts(YLt;2Lt)A80+Ts(YLt;2Rt)G74 (Roote et al., 1996)
Duplicated in	Dp(2;1)Scorv23 (Ashburner et al., 1983, Grau et al., 1984) Dp(2;2)TE35B-54L (Roote et al., 1996) In(2L)C163.41LTE35B-210R (Roote et al., 1996) In(2LR)Scorv1LTE35B-4R (Roote et al., 1996) In(2LR)Scorv1LTE35B-14R (Roote et al., 1996) Ts(2Lt;3Lt)G16+Ts(2Rt;3Rt)TE35B-3 (Roote et al., 1996) Ts(2Lt;4Lt)DTD22+Ts(2Rt;4Rt)TE35B-50 (Roote et al., 1996) Ts(YLt;2Lt)R15+Ts(Y;2Rt)el4 (Roote et al., 1996) Ts(YLt;2Lt)R15+Ts(Y;2Rt)TE35B-18 (Roote et al., 1996) Ts(YLt;2Lt)R15+Ts(YSt;2Rt)A80 (Roote et al., 1996)
Transgenic Constructs & Insertions
Transgenic Constructs
	Type of construct Name Expression data UAS construct P{GD1504} NA P{KK101464} NA P{NIG.3479R} NA
Insertions
	Type of insertions Name Expression data insertion of enhancer trap P{lacW}ospk13218 No P{PZ}osp00865 No P{PZ}osprJ571 No miscellaneous insertions Mi{MIC}ospMI00193 NA Mi{MIC}ospMI03036 NA Mi{MIC}ospMI05112 NA P{GT1}AdhBG01265 NA P{RS3}ospCB-5301-3 NA P{SUPor-P}ospKG06771 NA PBac{5HPw+}ospB307 NA PBac{PB}ospc00145 NA PBac{PB}ospc04765 NA PBac{PB}ospc05698 NA PBac{RB}ospe02984 NA PBac{RB}ospe02985 NA PBac{WH}ospf01299 NA PBac{WH}ospf03217 NA PBac{WH}ospf05495 NA PBac{WH}ospf05599 NA PBac{WH}ospf07101 NA insertion of enhancer trap binary system P{GawB}ospNP6350 No insertion of mobile activating element P{XP}ospd06776 NA P{XP}ospd11647 NA
Gene Ontology: Function, Process & Cellular Component ( 1 unique term )
	There is no information supporting a biological process or cellular component term for this gene. (Based on a review of available data before: 2006)
Terms Based on Experimental Evidence ( 0 terms )
Terms Based on Predictions or Assertions ( 1 term )
Molecular Function
CV term Evidence References
phospholipid binding inferred from electronic annotation with InterPro:IPR001849 (FlyBase Curators et al., 2004-)
Biological Process ( 0 terms)
Cellular Component ( 0 terms)
Sequence Ontology: Class of Gene
	nuclear_gene   protein_coding_gene
Interactions & Pathways
Summary of Physical Interactions
Summary of Genetic Interactions
Interacts with	Please look at the allele data for full details of the genetic interactions osp allele Gene References
External Data
Linkouts	BioGRID - A database of protein and genetic interactions • 60874 DroID - A comprehensive database of gene and protein interactions. • FBgn0003016 InterologFinder Protein-protein interactions (PPI) from both known and predicted PPI data sets. • 34850
Orthologs
OrthoDB Orthologs (37) - based on analysis using Dmel annotation version 5.41
OrthoDB Ortholog Groups
	OrthoDB orthology group searches (Waterhouse et al., 2011, Nucleic Acids Res. 39(Database issue): D283--D288) Drosophila inclusive ortholog search EOG6Z8ZFT Dipteran inclusive ortholog search EOG666VTW Insect inclusive ortholog search EOG615F8B Arthropod inclusive ortholog search EOG68PK14 Metazoa inclusive ortholog search EOG661GWV
Orthologs in Drosophila Species (EOG6Z8ZFT)
Organism Common Name Gene AAA Syntenic Ortholog Multiple Dmel Genes in this Orthologous Group Drosophila melanogaster fruit fly  Dmel\osp     Drosophila simulans   Dsim\GD21995 Y   Drosophila sechellia   Dsec\GM14739 Y   Drosophila erecta   Dere\GG24247 Y   Drosophila yakuba   Dyak\GE24875 Y   Drosophila ananassae   Dana\GF15007 Y   Drosophila pseudoobscura pseudoobscura   Dpse\GA17472 Y   Drosophila persimilis   Dper\GL25755 Y   Drosophila willistoni   Dwil\GK18083 Y   Drosophila virilis   Dvir\GJ18021 Y   Drosophila mojavensis   Dmoj\GI10322 Y   Drosophila grimshawi   Dgri\GH13845 Y
Orthologs in non-Drosophila Dipterans (EOG666VTW)
Organism Common Name Gene Multiple Dmel Genes in this Orthologous Group Aedes aegypti Yellow fever mosquito  Aaeg\AAEL008670   Anopheles gambiae Malaria mosquito  Agam\AGAP009847   Culex quinquefasciatus Southern house mosquito  Cqui\CPIJ009066
Orthologs in non-Dipteran Insects (EOG615F8B)
Organism Common Name Gene Multiple Dmel Genes in this Orthologous Group Apis mellifera Western honey bee  Amel\GB55755   Apis mellifera Western honey bee  Amel\GB48192   Acromyrmex echinatior Panamanian leafcutter ant  Aech\AECH14986   Atta cephalotes Leafcutter ant  Acep\ACEP17966   Camponotus floridanus Florida carpenter ant  Cflo\CFLO23469   Harpegnathos saltator Jerdons jumping ant  Hsal\HSAL12453   Linepithema humile Argentine ant  Lhum\LH19575   Pogonomyrmex barbatus Red harvester ant  Pbar\PB24349   Solenopsis invicta Red fire ant  Sinv\SINV18603   Acyrthosiphon pisum Pea aphid  Apim\ACYPI008555   Bombyx mori Silkmoth  Bmor\BGIBMGA006651   Pediculus humanus Human body louse  Phum\PHUM051580   Tribolium castaneum Red flour beetle  Tcas\TC013010   Tribolium castaneum Red flour beetle  Tcas\TC008527   Tribolium castaneum Red flour beetle  Tcas\TC006898
Orthologs in non-Insect Arthropods (EOG68PK14)
Organism Common Name Gene Multiple Dmel Genes in this Orthologous Group Daphnia pulex Water flea  Dpux\JGI_V11_318416   Daphnia pulex Water flea  Dpux\PCSK1N   Ixodes scapularis Deer tick  Isca\ISCW007121
Orthologs in non-Arthropod Metazoa (EOG661GWV)
Organism Common Name Gene Multiple Dmel Genes in this Orthologous Group Caenorhabditis elegans Nematode  Cele\F10G8.8   Strongylocentrotus purpuratus Purple sea urchin  Spur\SPU_023278gn   Strongylocentrotus purpuratus Purple sea urchin  Spur\SPU_022013gn   Strongylocentrotus purpuratus Purple sea urchin  Spur\SPU_026141gn
Human Orthologs (0)
Gene OMIM HGNC
AAA Orthologs (11) based on analysis using Dmel annotation version 4.3
Organism Gene Drosophila simulans Dsim\GD21995 Drosophila sechellia Dsec\GM14739 Drosophila erecta Dere\GG24247 Drosophila yakuba Dyak\GE24875 Drosophila ananassae Dana\GF15007 Drosophila pseudoobscura pseudoobscura Dpse\GA17472 Drosophila persimilis Dper\GL25755 Drosophila willistoni Dwil\GK18083 Drosophila virilis Dvir\GJ18021 Drosophila mojavensis Dmoj\GI10322 Drosophila grimshawi Dgri\GH13845
Stocks & Reagents
Stocks Listed in FlyBase ( 28 )
Bloomington	1023 osp1
Harvard	- P{XP}ospd06776 - P{XP}ospd11647 - PBac{PB}ospc00145 - PBac{PB}ospc04765 - PBac{PB}ospc05698 - PBac{RB}ospe02984 - PBac{RB}ospe02985 - PBac{WH}ospf01299 - PBac{WH}ospf03217 - PBac{WH}ospf05495 - PBac{WH}ospf07101
	More stocks available...
Genomic Clones ( 2 )
	BACR17M04 BACR32C03 Please Note FlyBase no longer curates genomic clone accessions so this list may not be complete
cDNA Clones ( 416 )
	Please Note This section lists cDNAs and ESTs that fall within the genomic extent of the gene model, which may include cDNAs and ESTs of genes within introns, or of overlapping genes. Please see GBrowse for alignment of the cDNAs and ESTs to the gene model.
cDNA Clones, Fully Sequenced
BDGP DGC clones	GH27591 LD14119 LD15891
Other clones	IP15972
cDNA Clones, End Sequenced (ESTs)
BDGP DGC clones	GH18414 GM21379 HL08134 LD13781 LD14764 LP10204 LP12918 LP19591 RE05714 RE11569 RE49627 RH04324 RH04390 RH07193 RH10576 RH10904 RH11852 RH12090 RH13479 RH13709 RH16019 RH16209 RH17093 RH17215 RH17293 RH17409 RH18095 RH20543 RH21653 RH21844 RH22177 RH22556 RH22735 RH26972 RH29611 RH29690 RH29717 RH30058 RH30182 RH30333 RH30767 RH31611 RH31616 RH32493 RH33778 RH37622 RH38076 RH39930 RH42014 RH42077 RH42543 RH42985 RH44978 RH45176 RH46016 RH46595 RH46639 RH47060 RH47229 RH47315 RH47775 RH48091 RH49746 RH50730 RH51215 RH51705 RH51995 RH52448 RH52603 RH54383 RH54653 RH55168 RH55173 RH55446 RH55941 RH56741 RH56809 RH56819 RH56839 RH56978 RH57786 RH58019 RH58044 RH58239 RH59131 RH60268 RH60713 RH61408 RH61433 RH61508 RH61691 RH62378 RH63266 RH64377 RH65659 RH66252 RH66273 RH66292 RH67049 RH67365 RH67395 RH67957 RH68761 RH68859 RH69793 RH71145 RH73083 RH74330 SD02265 SD09524 SD11290 SD16292 SD16426 SD18363 SD25304
Other clones	16646 19252 44607 52995 58257 59693 64086 66620 78246 79703 83874 97909 102711 108777 163447 214213.25 214345.22 214373.30 214375.34 277420 AI124308 CK01.14C.H6 CK01.36C.G12 CK01.61A.H4 CK01.81C.H4 CK01.99B.A10 CK01.104A.F2 CK01.104A.F4 CK01.105C.G5 CK01.124A.A8 CK01.142D.E7 CK00330 CK00365 CK01023 CK01258 CK01331 CK01336 CK01363 CK01421 CK01448 CK01456 CK01462 CK01603 CK01617 CK01634 CK01718 CK01792 CK01833 CK01958 CK02010 CK02026 CK02083 CK02089 CK02102 CK02134 CK02139 CK02216 CK02225 CK02311 CK02315 CK02322 CK02323 CK02429 CK02526 CK02564 CK02648 Dmel_dig_012356_0617_1329 Dmel_dig_013834_0625_2938 Dmel_neb_011377_0908_1005 Dmel_neb_016094_0595_2304 EC00884 EC41454 EK014948 EK027108 EK030002 EK031312 EK032025 EK034826 EK043120 EK047843 EK048304 EK049563 EK050409 EK053809 EK058252 EK058646 EK059016 EK063402 EK063762 EK064262 EK072843 EK073259 EK074745 EK075111 EK077604 EK082259 EK085812 EK087254 EK087859 EK088227 EK089024 EK089958 EK092228 EK095603 EK104936 EK105342 EK109250 EK113540 EK115509 EK116020 EK116527 EK117157 EK117938 EK119258 EK121312 EK127932 EK129929 EK130147 EK132358 EK133510 EK139758 EK145429 EK147049 EK150160 EK150812 EK154064 EK159214 EK160015 EK160129 EK161453 EK164214 EK165022 EK165624 EK165964 EK166151 EK170112 EK172611 EK178057 EK178259 EK180663 EK182949 EK183856 EK183952 EK184525 EK184711 EK187045 EK189615 EK194421 EK194453 EK194822 EK197709 EK198161 EK198534 EK201106 EK203155 EK205033 EK206320 EK206804 EK207149 EK209011 EK209843 EK211521 EK215732 EK217506 EK218354 EK219351 EK220628 EK221844 EK225650 EK225919 EK227244 EK227895 EK228028 EK235231 EK240954 EK241147 EK243327 EK243447 EK246762 EK253707 EK258226 EK261906 EK262115 EK264714 EK265613 EK267119 EK267342 EK267649 EK267760 EK278350 EK278462 EK289455 EK289458 EK290364 EK291554 EK292041 EK296753 EK301716 EK303642 EK304364 EK304527 EN01164 EN02064 EN15045 EP00315 EP01012 EP02341 EP04728 EP04754 EP05547 EP07649 EP08720 EP08834 EP08943 EP09905 EP10203 EP10234 EP10464 EP10540 EP11254 EP11446 EP11516 EP11550 EP11622 EP11914 EP12856 EP13133 EP13438 EP13602 EP14842 EP15443 EP15961 EP16650 EP17315 EP17909 EP18944 EP19157 EP19847 EP21002 EP21045 ESG0111d.B21.1_D10 FGH106G08 RP004076094 RP004092766 RP004100420 RP004110662 RP004116921 RP004145915 RP004147935 RP004148149 RP004153389 RP004192662 RP004196066 RP004197262 RP004200061 RP004203490 RP004209637 RP004212765 RP004214025 RP004214112 RP004219487 RP004223054 RP004227031 RP004233106 RP004235199 RP004238412 RP004244363 RP004247338 RP004253612 RP004254322 RP004258157 RP004259420 RP004297485 RP004351711 RP004357221 RP004391935 RP004427729 RP004430358 RP004435887 RP004438029 RP004443409 RP004450905 RP004461582 RP004469935 RP004475197 RP004478738 RP004479358 RP004513311 RP004514764 RP004519283 RP004530932 RP004539518 RP004540623 RP004549394 RP004549743 RP004550832 RP004551155
RNAi & Array Information
Linkouts	DRSC - Results from RNAi screens. • FBgn0003016 GenomeRNAi - GenomeRNAi – A database for cell-based and in vivo RNAi phenotypes and reagents • 34850
Antibody Information
Other Information
Discoverer
Etymology
Identification
Relationship to Other Genes
Source for database identity of	Source for identity of: osp CG3479 (Kodira, 1999.11)
Source for database merge of	Source for merge of: osp anon-WO02059370.75 (FlyBase, 1996-)
Additional comments	"BG:DS09219.1" is located within osp and is predicted to be transcribed in the same direction as osp. It may therefore be part of the osp gene, if osp has an alternative transcript that has not yet been identified as a cDNA. (Ashburner et al., 1999) Source for merge of osp anon-WO02059370.75 was sequence comparison (date:051113). (FlyBase, 1996-) Since "osp" is very near the 2L breakpoint of T(2;3)Mpe, the "Mpe1" phenotype may be due to a dominant "osp" allele. (Ashburner et al., 1981) Allelism of osp and alw assumed on basis of phenotype and map position (Ashburner).
Other Comments
	osp is not a vital gene. (Ashburner et al., 1999) Adh and Adhr are nested within osp and osp has a transcription unit greater than 74kb. (McNabb et al., 1996)
External Crossreferences & Linkouts
Sequence Crossreferences
	RefSeq (Transcripts) • NM_078843 NM_165089 NM_001144354 NM_001169504 RefSeq (Proteins) • NP_523567 NP_723879 NP_001137826 NP_001162975 DDBJ / EMBL / GenBank • AA141849 AA439264 AA439528 AA441103 AAA70209 AAA70211 AAF44880 AAF44881 AAF53402 AAM49881 AAN10878 ACL83032 ACZ94261 AE003410 AQ026050 AQ026134 AQ073307 AW942302 AX664501 AY118512 CAA77329 CZ466231 CZ469498 CZ470194 CZ475345 CZ477844 CZ480752 CZ480753 CZ480755 CZ485760 CZ485761 CZ488084 CZ488085 CZ488200 CZ488201 CZ489874 M17837 M19547 Z00030 Entrez Gene - A searchable database of RefSeq genes. • 34850 UniProtKB/Swiss-Prot • Q27421
Other Crossreferences
	InterPro domains - A database of protein families, domains, and functional sites • Pleckstrin homology domain (IPR001849) Pleckstrin homology-like domain (IPR011993) BDGP expression data - Patterns of gene expression in Drosophila embryogenesis • LD14119
Linkouts
	BioGRID - A database of protein and genetic interactions • 60874 DroID - A comprehensive database of gene and protein interactions. • FBgn0003016 DRSC - Results from RNAi screens. • FBgn0003016 FLIGHT - Cell culture data for RNAi and other high-throughput technologies • FBgn0003016 FlyAtlas - Adult expression by tissue, using Affymetrix Dros2 array • CG3479-RA FlyMine - Integrated genomics database for Drosophila, Anopheles, and C.elegans • FBgn0003016 GenomeRNAi - GenomeRNAi – A database for cell-based and in vivo RNAi phenotypes and reagents • 34850 InterologFinder Protein-protein interactions (PPI) from both known and predicted PPI data sets. • 34850 modMine - Data generated by the modENCODE project. • FBgn0003016
Synonyms & Secondary IDs ( 13 )
Reported As
Symbol Synonym	alw   anon-WO02059370.75   BG:DS01486.5 (Ashburner et al., 1999.10.12) BG:DS01486.7 (Ashburner et al., 1999.10.12, Ashburner et al., 1999) BG:DS07721.2 (Ashburner et al., 1999.10.12) CG3479 (Whitfield, 2000.5.19, Mohit et al., 2006, Keleman et al., 2009.8.5) l(2)rJ571   osp (Burnette et al., 2005, Moorman et al., 2006, Mohit et al., 2006, Aerts et al., 2007, Moschetti et al., 2004, Dietzl et al., 2007, Levis et al., 2011.12.16, Wasbrough et al., 2010, Reid et al., 2012, Japanese National Institute of Genetics, 2012.5.21) unnamed (Kreitman, 1995.1.26, Treisman and Rubin, 1996) v(2)rJ571 (Spradling, 1999.12.9)
Name Synonym	arc-like wing (Sturtevant, 1948) arclike wing   outspread (Moschetti et al., 2004, Reid et al., 2012)
Secondary FlyBase IDs
	FBgn0004947 FBgn0010673 FBgn0027874 FBgn0065995
References ( 89 )
Generate a list of	All references relating to this gene ( 89 )
List References by type	Research paper  ( 40 ) Supplementary material  ( 4 ) Review  ( 3 ) Personal communication to FlyBase  ( 31 ) FlyBase analysis  ( 4 ) DNA/RNA sequence record  ( 1 ) Stock list  ( 1 ) Curated genome annotation  ( 2 ) Patent  ( 1 ) Website  ( 1 ) Book  ( 1 )
Recent research papers ( 2 )
	Reid et al., 2012, Front. Cell. Infect. Microbiol. 2: 24 Identification of genetic modifiers of CagA-induced epithelial disruption in Drosophila. [FBrf0219288] Morozova et al., 2011, Genetics 187(4): 1193--1205 Transcriptional networks for alcohol sensitivity in Drosophila melanogaster. [FBrf0214377] Show all research papers ...
Recent reviews (0)
	All reviews listed in FlyBase were published before 2011 Show reviews ...