Eyes rough in varying degrees and somewhat bulging.
Affected by genetic modifiers. More extreme at 17-19 than at
22-25. Heterochromatin and sex affect expression so that X/0
> X/X > X/Y > X/X/Y; also enhanced by Df(2R)M41A10; spa
haplo-4s have an exaggerated phenotype. RK2.
Dominant rough-eye allele; more extreme than spaCat.
spaA/spapol shows extreme poliert phenotype; spaA/spaCat
Posterior third or half of eye of heterozygote
rough; facets irregular and fused. Homozygous lethal. Stocks
vary in expression, presumably because of genetic modifiers.
X/X and X/0 flies that are spaCat/spa show the bulging eyes
and roughening of spa and the posterior fused facets of
spaCat; X/X/Y and X/Y flies have only the spaCat phenotype.
spaCat/spapol has fusion of facets over entire surface of eye
and roughness in posterior region of eye. spaCat/4-sim is
wild type. spaCat/+ more extreme than spaCat/+/+ (Davis,
1969, Genetics 61: 577-94). RK2.
spae(lz): sparkling-enhancer of lozenge
Homozygote wild type in absence of lz; eyes strongly
roughened in presence of heterozygous lz3, lz34, lz36, or lzD.
Slight eye roughening when both spae(lz) and a lz allele are
heterozygous. spae(lz)/spapol and spae(lz)/spap65 have very
rough eyes but normal tarsal claws and spermathecae. RK3.
spap61: sparkling-poliert type
Eyes small, rough, and glazed. More extreme than
spapol or spap65. Nonpigmented tarsal claws. RK1.
Eyes somewhat reduced in size, rough and partially
glazed. More extreme than spapol but less so than spap61.
Tarsal claws unpigmented and possibly reduced; reminiscent of
certain lozenge mutants. Pulvilli and accessory female reproductive structures appear normal. Heterozygote with spapol
and spap61 has affected tarsal claws as well as rough eyes.
Heterozygote with spa has slightly roughened eyes at 25 but
markedly roughened eyes at 18; female somewhat more extreme
than male. Viability and fertility good. RK1.
Eyes rather small; surface smooth and glassy. During second day of pupal life, retinula cells withdraw from
other cells of eye disk. SEM studies show irregular disposition and morphology of ommatidial hairs as well as numerous
necrotic pits over surface of eye [Oster and Crang, 1972,
Trans. Am. Microsc. Soc. 91: 600-02 (fig.); Strum-Tegethoff
and Dicke, 1974, Theor. Appl. Genet. 44: 762-65]. ERG absent
[Grossfield, Handbook of Genetics (King, ed.). Plenum, New
York, pp. 679-702]. spapol/spaCat has extreme phenotype;
spapol/spa slightly more extreme than spa (Sturtevant, 1961,
DIS 35: 47). Homozygote has excellent viability and fertility. RK1.
Bristles reduced, somewhat variably. Trichogen
irregularly displaced and usually partly converted to socket
(Lees and Waddington, 1942, DIS 16: 70). sv/sv/sv triplo-4
nearly normal. sv haplo-4 extreme shaven (Schultz, 1935, Am.
Naturalist 69: 30-54). Expression depends on
temperature: excellent at 19, overlaps wild type at 25, and
entirely wild type at 30. RK2.
Resembles svn more than sv. Bristles frequently
reduced to stumps. RK2.
Edith M. Wallace, unpublished.
More extreme than svn. Thorax denuded over large
areas. Phenotype more severe than H2; bristleless sockets
found on adult integumentary derivatives of all imaginal
discs; nearly 100% penetrance on thorax. Shafts, where
present, often bent, twisted, or forked; up to 97% of bristle
organs on wing costa and distal leg segments fail to produce
normal shafts. Bracts present when bristle normally formed;
otherwise bracts missing (Tobler, Rothenbuhler, and Nothiger,
1973, Experientia 29: 370-71). Both sexes sterile. RK2.
Extremely short bristles. Viability excellent.
Trichogen irregularly displaced, becoming more or less converted into tormogen [Lees and Waddington, 1943, Proc. Roy.
Soc. (London), Ser. B, 131: 87-110 (fig.)]. Polarity of
microchaetae in vicinity of double sockets disrupted such that
they form a whorl around socket [Toney and Thompson, 1980,
Experientia 36: 644-45 (fig.)]. In triplo-4 svn/svn/svn, the
phenotype is more nearly normal than in diplo-4. RK1.