*In D. erecta and D. yakuba there is a shared pericentric inversion at the base of the B.C element (2L.2R in D. melanogaster). Thus relative to D. melanogaster the B and C elements are now mixed from telomere to centromere. The new order is B/C and C/B.
**In D. pseudoobscura, and D. persimilis the A and D elements are fused to form a metacentric X or first (1) chromosome from the X and autosomal 3L arms of D. melanogaster. In addition to this fusion there is a small pericentric inversion or transposition of material from the base of the A element to the base of the D element. It is unclear whether there is a reciprocal movement of D material into the base of A.
***In D. willistoni there is a fusion of the F element into the proximal end of the E element. Thus there is no dot chromosome in this species and the new order is E/F. Note that D. willistoni has a similar if not identical A/D fusion as that seen in D. pseudoobscura and D. persimilis. However it does not show the A => D inversion/transposition.