FB2025_05 , released December 11, 2025
FB2025_05 , released December 11, 2025
CV Term Report
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General Information
Term transmedullary neuron Tm9 ID (Ontology) FBbt:00003797 (Fly Anatomy)
Definition Transmedullary narrow field neuron that has most of its dendritic arborization in medulla layers M2 and M3, with some extensions into M1, M4 and M5 (Gao et al., 2008; Ting et al., 2014), and presynapses in M3 (Gao et al., 2008). Its axon follows the posterior edge of its cognate medulla column (Ting et al., 2014), then crosses in the second optic chiasma and terminates in lobula layer 1 (Fischbach and Dittrich, 1989; Ting et al., 2014), where it outputs to T5 neurons (Shinomiya et al., 2014). It is a cholinergic neuron (Gao et al., 2008; Shinomiya et al., 2014). It expresses the transcription factor sim (Ozel et al., 2021). There is usually one of these cells per optic column (Nern et al., 2025).[ FlyBase:FBrf0049410 FlyBase:FBrf0204652 FlyBase:FBrf0206213 FlyBase:FBrf0224196 FlyBase:FBrf0225094 FlyBase:FBrf0247762 FlyBase:FBrf0262545 ]
Also Known As "Tm9"
Comment Neurotransmitter was assessed using a cha-GAL4DBD driver (Gao et al., 2008), single-cell transcript profiling and immunohistochemistry (Shinomiya et al., 2014). Connectivity in the lobula was assessed by serial section electron microscopy of a single column (Shinomiya et al., 2014).
Links to External Ontologies
VFB:FBbt_00003797
Annotations
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Data Class Field Records
Alleles (FBal)  PHENOTYPE_MANIFEST_IN       3
Insertions (FBti)  EXPRESSION_STAGE_TISSUE_POSITION       6
Insertions (FBti)  PHENOTYPE_MANIFEST_IN       2
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 Full annotation statements 
Relevant FlyBase reports
 Alleles  Genes  Insertions Constructs
 transmedullary neuron Tm9 (all annotations which use CV term, excluding "NOT" statements)       8      3      5      8      8     16
     transmedullary neuron Tm9 | P-stage       3       --       --       --       2       --
Spanning Tree (Parents/Children)
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transmedullary neuron
 |__narrow field transmedullary neuron_____
cholinergic neuron                         |
 |__adult cholinergic neuron_______________|
adult neuron                               |
 |__adult cholinergic neuron_______________|
secondary neuron                           |
 |__Notch ON hemilineage secondary neuron__|
Notch ON hemilineage neuron                |
 |__Notch ON hemilineage secondary neuron__|
optic lobe narrow field columnar neuron    |
 |__narrow field transmedullary neuron_____|
                                           transmedullary neuron Tm9  50 rec.
                                            |__transmedullary neuron Tm9a 1 rec.
                                            |__transmedullary neuron Tm9b 1 rec.
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Is a narrow field transmedullary neuron
Notch ON hemilineage secondary neuron
adult cholinergic neuron
Develops from immature transmedullary neuron Tm9
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Synonyms
Secondary IDs
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FlyBrain_NDB:10163
VFB:FBbt_00003797