Hemizygous males usually have sex combs on the second and third legs, show a partial transformation of the posterior wing compartment towards the haltere and show a partial transformation of abdominal segment A4 towards A5, at both 25 and 29^oC. Hemizygous males rarely show loss of the humerus and occasionally show partial transformation of abdominal segment A1 towards A2 at 25^oC. At 29^oC, both these phenotypes are more common. The expressivity and penetrance of these homeotic transformation phenotypes are comparable in ph-d² and ph-d⁴ males.

Adults viable.

Homozygous female or hemizygous male flies often show a complete loss of the callus and bristles on the humerus. The prothoracic spiracle is unaffected. The second and third legs are often partially transformed towards the first leg. This transformation is restricted to the anterior compartment, and is particularly noticeable distally, where an extra sex comb appears. The first abdominal segment occasionally shows a partial transformation to a more posterior segment; dorsally, pigment and large bristles appear on the tergite, and ventrally, the sternite acquires bristles characteristic of the second or third abdominal segment. The fourth abdominal segment is often partially transformed to the fifth or sixth abdominal segment in males. Lethal over deficiencies that uncover the locus.