A mutated nuclear export signal which lacks export activity (sequence changed from LALKLAGLDI to LALKLAGADI) has been inserted at the third residue from the N-terminus of RanGAPSd.Tag:HA.
Flies carrying RanGapSd.T:Ivir\HA1,T:Ocun\nes-P12 show a high level of segregation distortion against target chromosomes Rspss.