2Lt - 21C8 | 60D1 - 59E1 | 40F - 59D4 | 40F - 21D1 | 60D2 - 2Rt

bk1 << S << Pin << bk4

When heterozygous with In(2R)bw^VK viability is increased.

Eye color like bw or pr, mottled with darker spots that deepen in red color with age. With st or v, has pale orange ground with dark orange spots. bw^V1/bw shows sharply reduced levels of transcript from both the bw⁺ allele in cis with the rearrangement and the bw allele on the homologous second chromosome (Henikoff, and Dreesen, 1989). Extra Y chromosome, as with other variegated browns, suppresses brown color, giving red eye sparsely speckled or splotched with darker spots. Amount of wild-type eye color also responds to fourth-chromosome constitution; 4/C(4) > C(4) = 4/4 > 4/0, with the first approaching wild type and the latter nearly bw (Lindsley and Rokop). Larval Malpighian tubules normal (Glass, Brehme). Generally lethal homozygous and in combination with other brown-Variegateds. heterozygous fully viable heterozygous fully fertile. Expression of bw^V1 suppressed by extra Y (Holm and Chovnick, 1976) or extra chromosome 4 (Lindsley). Double crossovers in 2L but not 2R fairly frequent. Single exchange in region 21D1-40F of 2L between In(2LR)bw^V1 and a normal sequence produces a recombinant carrying left end of normal chromosome 2, which is duplicated for 21A1-C8 and deficient for 60D2-F5. Heterozygote for this recombinant poorly viable, fertile, brown-Variegated, Minute and dwarf with pebbled arc wings; deficient for locus of M(2)60E. Reciprocal recombinant deficient for 21A1-C8 and duplicated for 60D2-F5; heterozygote poorly viable, fertile, bw⁺, Minute and giant; deficient for al and RpA2. Inversion heterozygote shows interchromosomal effect (Valentin, 1972, Hereditas 72: 243-54).