male lethal with normal X (Muller and Stone, 1930) XX females viable (Muller and Stone, 1930) XX females low fertility (Muller and Stone, 1930) males die later in the egg stage than Df(1)A1/Y males, without separation of the germ layers (Poulson, 1940, J. Exp. Zool. 83: 271-325). Triploid intersexes very low fertility (Pipkin, 1940, Texas Univ. Publ. 4032: 126-56).