Approximately 40% of gonads of progeny of homozygous females agametic. Although some eggs of homozygous females exhibit abnormal polar granules, normal numbers of pole cells form; some pole cells abnormal with degenerating polar granules and nuclear bodies, but pole cells reach gonads at 14 hr of development and then in 40% of the gonads become necrotic and disappear; responses of right and left gonads correlated. Phenotype most pronounced at 25oC, decreasing at higher and lower temperatures. Expression in progeny of heterozygous females half that in those of homozygotes.