The mitotic divisions of the gonial cells and meiosis are completed in hemizygous males. The development of the axonemal sheath and its association with the mitochondrial derivatives is abnormal in midstage spermatids. Many mitochondrial derivatives are associated with cytoplasmic membranes and in some cases paracrystalline formation is induced at more than one position within a single mitochondrion. Paracrystalline formation is sometimes induced in both mitochondrial derivatives that are associated with an axoneme. Spermatid maturation proceeds through the elongation and individualisation stages in some cysts, although there are subtle abnormalities in individualisation and alignment. Spermatid heads condense in some cysts, but coiling is not completed and sperm do not escape into the seminal vesicles.

Involved in reproductive tissue development. Mitochondrial derivatives and axoneme atypical during spermatid development.

male-sterile