sv^spa-A heterozygotes have a similar, but less extreme, phenotype to sv^spa-1 homozygotes raised at 18^oC, with defects largely being restricted to the posterior portion of the eye. Some ommatidia in adults have an abnormal number of photoreceptor cells. At the early pupal stage, cone cells in ommatidia in the posterior of the retina frequently fail to assemble in the typical rhomboid-like configuration. After completion of ommatidial assembly, one or two cone cells or one of the primary pigment cells may be missing. In the latter case, a single primary pigment cell expands beyond its normal boundaries and attempts to encircle all four cone cells.

Rough eye phenotype. Heterozygotes with sv^spa-Cat are lethal and heterozygotes with sv^spa-pol have an extreme poliert phenotype.

RK1.

Dominant rough-eye allele; more extreme than sv^spa-Cat. sv^spa-A/sv^spa-pol shows extreme poliert phenotype; sv^spa-A/sv^spa-Cat lethal. RK1A.