Approximately 50% of homozygotes die as larvae, and 50% die as pharate adults with an eyeless phenotype. The pharate adults break open the operculum but usually fail to eclose. The few that do eclose remain attached by the wings to the pupal case and live for as long as 6 days. They show uncoordinated movement, fall over and flail around wildly. Homozygotes show massive head defects including the lack of one or both eyes, duplications and triplications of antennae, and greatly increased numbers of bristles in the postorbital, postgenal and antennal areas. Other defects include missing or swollen leg segments, increased leg bristle numbers, and poorly developed and non-expanded wings. Eye-antennal imaginal discs from homozygous larvae show abnormalities including duplications and triplications of the antennal discs, and reduced eye discs and optic lobe connections. Lethality is not temperature sensitive, but head and leg defects are more severe at lower temperatures.