Eggs and embryos produced by homozygous females are completely dorsalised. The dorsal appendages are enlarged and completely encircle the anterior end of the egg. Less than 1% of the eggs are fertilised. The egg chamber in homozygous females first appears abnormal at stage 10, when all the follicle cells surrounding the anterior end of the oocyte begin to thicken and migrate centripetally. This gives the oocyte a 'bomb-like' appearance. The egg chamber is also shorter than normal. fs(1)K1013 heterozygous or homozygous females also homozygous for grk3, grk6, cniAA112 or Egfrt1 produce eggs with a ventralised phenotype. fs(1)K1013 homozygous females also heterozygous for Egfrt1 or cniAA112 produce eggs with a dorsalised phenotype. fs(1)K1013 homozygous females also heterozygous for grk3 produce eggs with a range of phenotypes, from that of fs(1)K1013 homozygotes to almost wild-type. Approximately 70% of the eggs produce an intermediate phenotype with two dorsal appendages that are located at the very anterior end of the egg. The embryos derived from these eggs also show a range of phenotypes, from a dorsalised to an almost wild-type cuticle. Rescue always occurs first at the posterior end of the embryo, with the cephalic structures being the last to differentiate normally and often being everted.