Partial excision of the P{PZ} element, with 38bp of the P{PZ} sequences remaining. These sequences include two 15bp inverted repeats (that are vestiges of the 31bp inverted repeats of the parental P{PZ} element) and the 8bp target site duplication that accompanied the insertion of the parental P{PZ} element.
Three classes of put allele can be identified with respect to decreasing severity of wing venation phenotypes with put10 : Class I (put10460 = put62) > Class II (put51 = put88 = put135) > Class III (put10 = put24 = put97).