Malpighian tubules are short with fewer but larger cells than normal. The hindgut and foregut are short. Midgut constrictions do not form. Head involution is incomplete, tracheae are abnormal and there are holes in the epidermis.
thrk01302 was identified in the 'l(2)k01302' line. A deficiency uncovering the site of the P{lacW}tkvk01302 insertion complements the 'l(2)k01302' chromosome for lethality and does not produce embryos with the defects characteristic of the homozygous 'l(2)k01302' line. This suggests that the thrk01302 mutation (representing the lethality and embryonic defects) is not caused by the P{lacW}l(2)k01302k01302 insertion.