shotV104/shotA405 embryos have an excess of epidermal muscle attachment cells at a number of sites in the epidermis. The embryos have a significant disruption of the muscle pattern; in many cases individual myotubes are not oriented correctly and in some cases the myotubes round up. shotA405/Df(2R)MK1 embryos have an excess of epidermal muscle attachment cells at a number of sites in the epidermis.