Excision of the P{EP} element resulting in nucleotide exchanges and insertion of two nucleotides just upstream of the putative TATA box (the sequence CCCATCTA which occurs starting at position +72 relative to the P{EP}EP487 insertion site has been changed to CAAATC).
A few homozygous escapers are seen if the flies are grown at low density. The bristles on the notum of these adults, particularly in the anterior portion, show defects in their polarity. The flies also have a rough eye phenotype, which is due to the failure to form a regular array of hexagonal corneal lenses, which is accompanied by an irregular distribution of interommatidial bristles. The degree of rotation of the ommatidia appears random; some ommatidia have not rotated at all, while others have rotated only 45o or have continued the rotation beyond the normal 90o. The overall composition of photoreceptors in the ommatidia is normal.
