In contrast to the wild-type, in mutant larvae, the lamina plexus is discontinuous. Thick bundles of axons are frequently observed between the lamina and the medulla, in striking contrast to the thin bundles in wild-type. In the most severe cases, the lamina plexus is largely missing and the medulla neuropil is hyperinnervated. In addition, disruptions in the normally smooth topographic map in then medulla are observed. However R-cell projections do not extend through the medulla into deeper layers of the optic lobe, suggesting that R8 neurons terminate, as in wild-type, within the medulla neuropil.