Nucleotide substitution: C338A. Nucleotide substitution: G368A. Nucleotide substitution: T433C. Nucleotide substitution: C2041T. Nucleotide substitution: T2588A. Nucleotide substitution: A2629G. Nucleotide substitution: G2803A. 5bp deletion of CTTTT starting at nucleotide position 325. Also lacks the 35bp insertion starting at nucleotide position 392 which is present in the "S26-F" strain. (Substitutions are relative to the "S26-F" strain. Transcription start site corresponds to nucleotide position 1184).