FB2026_03 , released September 17, 2026
FB2026_03 , released September 17, 2026
CV Term Report
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General Information
Term fan-shaped body ID (Ontology) FBbt:00003679 (Fly Anatomy)
Definition The largest synaptic neuropil domain of the adult central complex, located posterior to the ellipsoid body and anterior to the protocerebral bridge (Hanesch et al., 1989; Wolff et al., 2015). The fan-shaped body is composed of a rough 2-dimensional grid of layers and columns (Hanesch et al., 1989; Wolff et al., 2015; Hulse et al., 2021). Eight horizontal layers, numbered bottom to top, form a fan shape and can be divided into vertical columns (sometimes called staves, slices or segments) (Ito et al., 2014; Wolff et al., 2015). An additional, much narrower, 9th layer, which lacks a columnar organization, sits dorsal to layer 8 (Wolff et al., 2015). The number of columns varies for different neuronal types (Scheffer et al., 2020).[ FlyBase:FBrf0049409 FlyBase:FBrf0212649 FlyBase:FBrf0221742 FlyBase:FBrf0224194 FlyBase:FBrf0227801 FlyBase:FBrf0246888 FlyBase:FBrf0254510 ]
Also Known As "fan shaped body" ; "FB" ; "fb"
Comment Previously thought to be 8 slices per hemisphere numbered from 1-8 medial to lateral - arranged in 4 closely associated pairs (Ito et al., 2014). Each of the two neighboring slices (1-2, 3-4, 5-6, 7-8) are associated more closely because they receive small-field columnar neurons generated by the same neuroblasts, forming four groups on each side of the midline, from lateral to medial: segment pair W, X, Y and Z (Boyan and Williams, 2011; Ito and Awasaki, 2008). Six to nine layers can be identified, depending on the staining that is used (Hanesch et al., 1989; Young and Armstrong, 2010; Kahsai and Winther, 2011, Lin et al., 2013, Wolff et al., 2015). Recent papers divide the FB into 9 layers (Wolff et al., 2015; Scheffer et al., 2020).
Links to External Ontologies
VFB:FBbt_00003679
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Data ClassFieldDirect annotations
Alleles (FBal)PHENOTYPE_MANIFEST_IN
Insertions (FBti)EXPRESSION_STAGE_TISSUE_POSITION
Insertions (FBti)PHENOTYPE_MANIFEST_IN
Constructs (FBtp)PHENOTYPE_MANIFEST_IN
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 Alleles Split System Combinations Genes Insertions Constructs
 fan-shaped body (all annotations which use CV term, excluding "NOT" statements)
     fan-shaped body | adult stage
      --       --       --
     fan-shaped body | conditional
      --       --       --
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adult central complex
 |__central body________________
multi-cell-component structure  |
 |__synaptic neuropil domain____|
synaptic neuropil               |
 |__synaptic neuropil domain____|
synaptic neuropil domain        |
 |__central body________________|
                                fan-shaped body  1834 rec.
                                 |__adult dorsal fan-shaped body 15 rec.
                                 |   |__fan-shaped body layer 6 3 rec.
                                 |   |__fan-shaped body layer 7 2 rec.
                                 |   |__fan-shaped body layer 8 7 rec.
                                 |   |__fan-shaped body layer 9
                                 |__adult ventral fan-shaped body 7 rec.
                                 |   |__fan-shaped body layer 1(+) 7 rec.
                                 |   |__fan-shaped body layer 2 1 rec.
                                 |   |__fan-shaped body layer 3 3 rec.
                                 |   |__fan-shaped body layer 4 3 rec.
                                 |   |__fan-shaped body layer 5 7 rec.
                                 |__fan-shaped body column
                                 |__fan-shaped body layer 33 rec.
                                     |__fan-shaped body layer 1(+) 7 rec.
                                     |__fan-shaped body layer 2 1 rec.
                                     |__fan-shaped body layer 3 3 rec.
                                     |__fan-shaped body layer 4 3 rec.
                                     |__fan-shaped body layer 5 7 rec.
                                     |__fan-shaped body layer 6 3 rec.
                                     |__fan-shaped body layer 7 2 rec.
                                     |__fan-shaped body layer 8 7 rec.
                                     |__fan-shaped body layer 9
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Is a synaptic neuropil domain
Develops from embryonic/larval fan-shaped body - primordial
Part of central body
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VFB:FBbt_00003679