FB2026_03 , released September 17, 2026
CV Term Report
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Term medulla layer M4 ID (Ontology) FBbt:00003753 (Fly Anatomy)
Definition Fourth most lateral layer of the medulla. It corresponds to the narrow layer between the proximal end of lamina neuron L3 and the distal border of lamina neuron L1's proximal arborization (Takemura et al., 2008).[ FlyBase:FBrf0049410 FlyBase:FBrf0205531 FlyBase:FBrf0224194 ]
Also Known As"L4 layer" ; "medulla layer 4" ; "stratum M4"
CommentIn the literature there are two methods for describing spatial location of neurons in the visual system: distal-proximal (e.g. Fischbach and Dittrich, 1989), and medial-lateral (e.g. Ito et al., 2014), both meaning outer and inner, and refer to the distance of a structure along the visual pathway relative to the center of the brain. We use the classical 'distal-proximal'.
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VFB:FBbt_00003753
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Insertions (FBti)EXPRESSION_STAGE_TISSUE_POSITION
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medulla
 |__medulla layer________
synaptic neuropil layer  |
 |__medulla layer________|
plexiform medulla        |
 |__outer medulla________|
medulla layer            |
 |__outer medulla________|
                         medulla layer M4  4 rec.
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VFB:FBbt_00003753