FB2026_03 , released September 17, 2026
FB2026_03 , released September 17, 2026
CV Term Report
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General Information
Term distal medullary amacrine neuron Dm4 ID (Ontology) FBbt:00003771 (Fly Anatomy)
Definition Distal medullary wide-field amacrine neuron whose cell body is located in the anterior region of the cell body rind of the medulla. The main neurite extends into M6, and then turns back to arborize in the boundary between M5 and M6, and in M3B, with mixed morphology arborizations. The arbor varies in shape between cells, covering around 21 columns, but with these areas tiling. It is located in a central position in the column, surrounded by the terminal of photoreceptors R7 and R8, with the cell boundaries following the column boundaries. There are around 40 Dm4 neurons per hemisphere. They are glutamatergic (Davis et al., 2020).[ FlyBase:FBrf0049410 FlyBase:FBrf0204652 FlyBase:FBrf0228639 FlyBase:FBrf0244935 ]
Also Known As "Dm4"
Comment Classified as columnar based on first stretch of main projection being limited to a single column, but as it then bends and projects along the plane of M5/M6 where it arborizes, this is clearly an edge case. Pre- versus postsynaptic innervation were judged by scoring of terminal morphology from figures in Fischbach and Dittrich (1989) as assessed by FlyBrain Neuron DB.
Links to External Ontologies
VFB:FBbt_00003771
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Data ClassFieldDirect annotations
Alleles (FBal)PHENOTYPE_MANIFEST_IN
Insertions (FBti)EXPRESSION_STAGE_TISSUE_POSITION
Insertions (FBti)PHENOTYPE_MANIFEST_IN
Constructs (FBtp)PHENOTYPE_MANIFEST_IN
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 distal medullary amacrine neuron Dm4 (all annotations which use CV term, excluding "NOT" statements)
     distal medullary amacrine neuron Dm4 | adult stage | decreased number
      --       --
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     distal medullary amacrine neuron Dm4 | adult stage | increased number
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adult optic lobe neuron
 |__distal medullary amacrine neuron________
optic lobe amacrine neuron                  |
 |__distal medullary amacrine neuron________|
adult optic lobe columnar neuron            |
 |__optic lobe wide-field columnar neuron___|
adult neuron                                |
 |__adult glutamatergic neuron______________|
secondary neuron                            |
 |__Notch OFF hemilineage secondary neuron__|
Notch OFF hemilineage neuron                |
 |__Notch OFF hemilineage secondary neuron__|
medulla intrinsic neuron                    |
 |__distal medullary amacrine neuron________|
glutamatergic neuron                        |
 |__adult glutamatergic neuron______________|
                                            distal medullary amacrine neuron Dm4  32 rec.
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Is a distal medullary amacrine neuron
Notch OFF hemilineage secondary neuron
adult glutamatergic neuron
optic lobe wide-field columnar neuron
Develops from immature distal medullary amacrine neuron Dm4
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FlyBrain_NDB:10084
VFB:FBbt_00003771