bk1 << ac << rb << bk2 bk3 << fu << car << bk4
Heterozygous female has stubby posterior verticals and disarranged scutellars; outer wing margin less convex than normal; fair viability and fertility. Enhances expression of heterozygous B. male lethal.
Duplicated for 1B1-4D2 and 18A4-20F.