Imprecise excision of the progenitor insertion, resulting in a 2225bp deletion extending from the original insertion site. The deletion encompasses the entire intergenic region between Drep-2 and Mad1 and the first 814bp of the Mad1 coding sequence (removing the first 272 of 730 codons of Mad1). The first transcription start site of Drep-2 is also removed.
Separable from: a linked mutation outside the genomic region delimited by Df(2R)45-30n that causes male sterility.