Ovarian tumours, ovaries contain many egg chambers filled with small cells; follicle cells secrete a tiny round egg shell around the egg chambers.
Homozygous females contain many small egg chambers in their ovaries, which seem blocked in early stages of oogenesis; follicle cells nevertheless synthesize a very small, round chorion around these egg chambers, giving them the appearance of small round peppercorns.
Schupbach and Wieschaus.