Heterozygotes often show weak transformation of the second and third legs to first legs, and partial transformation of the fourth abdominal segment to fifth abdominal segment. Pcl¹/Pcl³ flies usually die as pharate adults. Head structures usually appear normal, although, rarely, outgrowths of leg tissue (as seen by the presence of bracted bristles) occur on the antennae. The prothorax, including the first leg, and humerus, mesonotum and dorsal metathorax appear normal. Wing vein L5 often fails to reach the wing margin, possibly indicating a weak transformation of wing to haltere. Almost the entire anterior compartment of the second leg is transformed into a first leg anterior compartment in most cases. The posterior compartment of the second leg contains a mixture of first, second and third leg posterior structures. Transformation of the third leg to first leg occurs mainly in the anterior compartment. This transformation is complete distally, with sex combs of 9-13 teeth forming in the basitarsus, but is incomplete proximally, with the coxa being only slightly affected. The posterior compartment of the third leg appears almost normal, although there is some indication of transformation to first or second leg; occasionally a bristle forms in the central region of the trochanter posterior compartment, small groups of trichomes often occur in the central portion of the femur posterior compartment, and the bristles at the ventral edge of this region are slightly enlarged. All the abdominal tergites of Pcl¹/Pcl³ flies show variegated transformation towards more posterior segments. The 1st abdominal tergite often contains patches of cuticle that are transformed to 2nd, 3rd or perhaps 4th tergite. The 2nd, 3rd and 4th tergites often show patches of cuticle that are transformed to 5th or 6th tergite. The 5th tergite is usually transformed to 6th tergite, and the 6th tergite is usually transformed to 7th tergite, which results in the loss of the 6th tergite in males. The external genitalia and analia are normal in Pcl¹/Pcl³ flies. Internally, females show transformation of the parovaria into spermathecae. The internal genitalia of males are normal.

homozygous viable; leg2, leg3 transformed to leg1 (het)