Homozygotes strongly semi-lethal in males; weakly so in females; homozygous females sterile; produce no eggs. Homozygous males have reduced fertility. Survival sensitive to the presence of the Y chromosome as well as to the presence of 0.1 M butyrate in the medium. homozygotes butyrate sensitive; homozygous semi-lethal homozygous sterile
Isolated as a suppressor of position effect variegation at the w locus in flies carrying In(1)wm4h, and a dominant enhancer of position effect variegation, E(var)c1011.