Frameshift after amino acid residue 1693 due to a deletion of 4bp.
A four base pair deletion leads to a frame shift.
Eyes roughened due to abnormalities in the hexagonal facet array: most facets include 1-3 extra outer photoreceptor cells which are probably mystery cells. Ovaries of homozygous females are normal in appearance but the embryos produced never form syncytial blastoderm and all fail to hatch.