Transheterozygotes are female sterile and semi-lethal, exhibit rough eyes, bristle and wing defects (both enhanced at 18oC), brain defects, meiosis is mostly normal with some nondisjunction. Heterozygotes with Df(3R)Hdγ1 are viable but female sterile.
Phenotype assayed in trans with a multiple mutant chromosome carrying mutations of polo, mgr, asp, stg and gnu.