Splice donor site of the second intron has been mutated from GTATG to ATATG (G to T mutation at coordinate 2R:12570993 ).
Females have rudimentary ovaries that either lack germ cells, or contain a few egg chambers with abnormal numbers of germ cells. Follicle cells function normally. Posterior migration of the follicle cells occurs normally but they secrete reduced dorsal appendages and an 'open-ended' chorion.