The germ band twists laterally, rather than extending dorsally as in wild type. Proper orientation of the germ band in recovered by the completion of germ-band extension. Embryos exhibit mislocalisation of the pericardial cells and there are fewer of these cells than in wild type. Embryos exhibit significant gaps in the dorsal trunk of the trachea. Embryos frequently display one salivary gland misshapen and smaller than the other, the gland may also be shifted closer to the midline.