Insertion of a P{GS} element in the 3' flanking region.
Flies expressing P{GS}exuGS2220 under the control of Scer\GAL4c355 show semi-lethality and have fused wing veins. Flies expressing P{GS}exuGS2220 under the control of Scer\GAL4dpp.blk1 have fused tarsal segments.