Mutants show significantly lower memory than control flies at multiple times after olfactory classical conditioning.

In odour avoidance tests, mutant flies avoidance the aversive odours of octanol and benzaldehyde to the same degree as wild-type controls. Odour response after preshock is also wild-type. In odour learning tests, mutant flies exhibit relatively poor performance compared to wild-type flies when tested 3 minutes after odour conditioning. When trained with a short program schedule, mutants perform poorly relative to wild-type, but after normalisation, memory is seen to decay at an identical rate. Memory retrieval also appear to be wild-type. The defect appears to be in memory formation.