Homozygotes have rough eyes and often have nicks in the wing margin. Homozygous females lay eggs with slightly shorter dorsal appendages than normal. The dorsal-ventral chorion pattern of these eggs and the position of the dorsal appendages is normal.
E(To)99 is an enhancer of some die during embryonic stage | maternal effect | recessive phenotype of Tm1el4/Tm1eg9, osk1
E(To)9[+]/E(To)99 is an enhancer of embryonic abdominal segment phenotype of Tm1el4/Tm1eg9, osk1
E(To)9[+]/E(To)99 is an enhancer of embryonic abdominal segment phenotype of Tm1el4/Tm1eg9, osk[+]/osk1