Eggs derived from homozygous mutant germline clones do not undergo nuclear divisions.

Spindle formation in meiosis I appears normal in homozygous oocytes. In homozygous eggs, the first meiotic spindle is anastral (as in wild type) and appears to function properly. However, meiosis II is severely disrupted in these eggs; the central array of microtubules is absent, and instead two anastral spindles are formed, which are not properly aligned to each other and to the cortex, or the spindles are strongly disorganised.