Imprecise excision of the progenitor insertion.
microtubule & oocyte
The retained eggs laid by homozygous females have many morphological defects; many have fused dorsal appendages, many are smaller than wild type, and some have open anterior chorions.
A zone along the anterior region of the oocyte shows a significantly higher concentration of microtubules in homozygous egg chambers compared to wild type.
tral[+]/tral1Δ1 is an enhancer of dorsal appendage | maternal effect phenotype of BicC4